All variations which are demonstrably useful can be similarly dealt
with if their origin is explained by variational tendencies.
We perceive that the objection which Von Hartmann brings against
heredity is only valid on the ground that this process affords no
security for the preservation of variations which occur singly. That
heredity itself is a mechanical process is not directly disputed; it is
simply assumed that new characters can be transferred by inheritance
only when they are produced by the metaphysical “developmental
principle,” and not when they arise “accidentally.” This critic does
not therefore direct his attack against heredity, but rather against
the mechanical origin of variability.
Von Hartmann might have said here that a reference of the phenomenon
of heredity to purely mechanical causes, _i.e._ a mechanical theory
of heredity, is up to the present time wanting. That he has not done
so proves on the one hand that he despised the dialectical art, but,
on the other hand, that he himself has not overlooked the subserviency
of the total phenomenon to law, and that he grants the possibility of
finding a mechanical explanation therefor. If, in fact, the power of
inheritance does not depend upon mechanical principles, I know not
what organic processes we are entitled to regard as mechanical, since
they are all dependent in essence upon heredity, with which process
they are at one, and from which they cannot be thought of as isolated.
Haeckel correctly designates reproduction as surplus individual
growth, and accordingly refers the phenomena of heredity to those of
growth. Conversely, growth may also be designated reproduction, since
it depends upon a continuous process of multiplication of the cells
composing the organism, from the germ-cell to the innumerable congeries
of variously differentiated cells of the highly developed animal body.
Who can fail to see that these two processes, the reproduction of the
germ-cell and its offspring in the economy of the individual, and the
reproduction of individuals and species in the economy of the organic
world, show an exact and by no means simply superficial analogy?[296]
But whoso grants this must also conceive both processes to depend upon
the same cause--he cannot assume for the one a causal power and for
the other a directive principle. If nutrition and cell-multiplication
are purely mechanical processes, so also is heredity. Although it
has not yet been possible to demonstrate the mechanism of this
phenomenon, it can nevertheless be seen broadly that by means of a
minimum of living organic matter (_e.g._ the protoplasm of the sperm
and germ-cell) certain motions are transferred, and these can be
regarded as directions of development, as I have already briefly laid
down in a former work.[297] The power of organisms to transmit their
properties to their offspring appears to me to be only conceivable in
such a manner “that the germ of the organism by its chemico-physical
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