composition together with its molecular structure, has communicated to
it a fixed direction of development--the same direction of development
as that originally possessed by the parental organism....” (_loc.
cit._ p. 24). This is confessedly nothing more than a hint, and we do
not learn therefrom the means by which developmental direction can be
possibly transferred to another organism.
Recently Haeckel, that indefatigable pioneer to whom we are indebted
for such a rich store of new ideas, has attempted to bridge over this
gap in his essay on “The Perigenesis of the Plastidule,” Berlin, 1876.
The basic idea, that heredity depends upon the transference of motion,
and variability upon a change of this motion, completely corresponds
with the conviction gained in the province of physical science, that
“all laws must finally be merged in laws of motion” (Helmholtz[298]).
I hold this view to be the more completely justifiable--although
certainly not in the remotest degree as proved--because I formerly
designated the acquired individual variations as the “diversion of the
inherited direction of development.” Haeckel’s hypothesis in so far
accomplishes more than Darwin’s pangenesis, in which a transference
of matter, and not of a species of motion peculiar to this matter, is
assumed. But although the germ of a mechanical theory of heredity may
be contained in Haeckel’s hypothesis, this nevertheless appears to me
to be somewhat remote from completely solving the problem. It brings
well into prominence one portion of the process of inheritance; under
the image of a molecular motion of the plastidule, which motion is
modifiable by external influences, we can well understand the fact of
a change gradually taking place in the course of generations. On the
other hand, the assumption of consciousness in the plastidule,--however
admissible philosophically--although only as a formula, scarcely
furnishes any deeper knowledge. In the light of a theory, detailed
instances which were formerly obscure should become comprehensible.
I fail to see, however, how the various forms of atavism, _e.g._ the
reversions which so commonly occur by crossing different races, become
more comprehensible by assuming consciousness in the plastidule. If
in both parents the plastidule long ago acquired different molecular
motions, why, in its rencounters in the germ, does it recollect past
times and reassume the older and long abandoned motion? That it does
acquire the latter is indeed a fact if we once refer the directional
development of the individual to molecular motion of the plastidule;
the wherefore does not appear to me, however, to become clearer by
assuming consciousness in the plastidule. A mechanical theory of
heredity must rather be able to show that the plastidule movements of
the male and female germ-cells, in their rencounter in the case of the
crossing of widely divergent forms, become mutually modified in such
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