An especially striking instance of incongruence is offered by the
two species of _Chærocampa_ most common with us, viz., _Elpenor_ and
_Porcellus_, the large and small Elephant Hawk-moths. The larvæ are
so similar, even in the smallest details of marking, that they could
scarcely be identified with certainty were it not that one species
(_Elpenor_) is considerably larger and possesses a less curved caudal
horn than the other. The moths of these two species much resemble
one another in their dull green and red colours, but differ in the
arrangement of these colours, _i.e._ in marking, and also in the form
of their wings, to such an extent that _Porcellus_ has been referred to
the genus _Pergesa_[193] of Walker. If systemy, as is admitted on many
sides, has only to indicate the morphological relationship, this author
is not to blame--but in this case a special larval classification must
likewise be admitted, in a manner somewhat similar to that at present
adopted provisionally in text-books of zoology for the Hydroid Polypes
and inferior Medusæ. This case of _Porcellus_, however, shows that
those are correct who maintain that systemy claims to express, although
incompletely, the blood-relationship, and that systematists have always
unconsciously formed their groups as though they intended to express
the genetic connection of the forms. Only on this supposition can it
appear incorrect to us to thus separate two species of which the larvæ
agree so completely.
I cannot conclude this review of the various systematic groups without
taking a glance at the groups comprised within species, viz. varieties.
Whilst in species incongruence is of frequent occurrence, in varieties
this is the rule, for which reason it admits in this case of being more
sharply defined, since we are not concerned with a double difference
but only with the question whether in the one stage a difference or an
absolute similarity is observable. By far the majority of varieties are
either simply imaginal or merely larval varieties--only the one stage
diverges, the other is quite constant.
Thus, as has already been shown, in all the seasonally dimorphic
butterflies known to me the caterpillars of the two generations of
imagines, which are often so widely different, are exactly alike; and
the same obtains for the majority of purely climatic varieties of
butterflies. Unfortunately there are as yet no connected observations
on this point. The only certain instance that I can here mention is
that of the Alpine and Polar form of _Pteris Napi_. This variety,
_Bryoniæ_, the female of which differs so greatly in marking and
colouring, possesses larvæ which cannot be distinguished from those of
the ordinary form of _Napi_.(See part I. appendix I. p. 124.)
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