The explanation of the phenomena thus far lies on the surface, and it
is scarcely possible to advance any other; but why should one stage
become changed more frequently or to a greater extent than the other?
why should one portion be induced to change more frequently or more
strongly than another? whence come these inducements to change? These
questions bring us to the main point of inquiry:--Are the causes which
give rise to these changes internal or external? Are the latter the
result of a phyletic vital force, or are they only due to the action of
the external conditions of life?
Although an answer to this question will be found in the preceding
essay, I will not support myself on the results there obtained, but
will endeavour to give another solution of the problem on fresh
grounds. The answer will indeed be the same as before:--A phyletic
force must be discountenanced, since in the first place it does not
explain the phenomena, and in the second place the phenomena can be
well explained without its assumption.
The admission of a phyletic vital force does not explain the phenomena.
The assumption that there is a transforming power innate in the
organism indeed agrees quite well with the phenomenon of congruence,
but not with that of incongruence. Since a large number of cases of
the latter depend upon the fact that the larvæ are more frequently
influenced by causes of change than their imagines, or _vice versâ_,
how can this be reconciled with such an internal force? On this
assumption would not each stage of a species be compelled to change, if
not contemporaneously at least successively, with the same frequency
and intensity, by the action of an innate force? and how by means of
the latter can there ever result a greater form-divergence in the larvæ
than in the imagines?
It is delusive to believe that these unequal deviations can be
explained by assuming that the phyletic force acts periodically.
Granting that it does so, and that the internal power successively
compels the imago, pupa, and finally the larva to change, there would
then pass a kind of wave of transformation over the different stages
of the species, as was actually shown above to be the case in the
single larval stages. The only possible way of explaining the unequal
distances between larvæ and imagines would therefore be to assume that
two allied groups, _e.g._ species, were not contemporaneously affected
by the wave, so that at a certain period of time the imago alone of one
species had become changed, whilst in the other species the wave of
transformation had also reached the larva. In this case the imagines of
the two species would thus appear to be more nearly related than their
larvæ.
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