But if this is the case--if within the period of time which is
sufficient for the production of species, the one stage only is but
seldom and quite exceptionally influenced by transforming impulses,
whilst both stages are as a rule affected, although not with the same
frequency, it must necessarily follow that on the whole, as the period
of time increases, the difference in the number of these impulses which
affect the larva and of those which affect the imago must continually
decrease, and with this difference the magnitude of the morphological
differences resulting from the transforming influences must at the same
time also diminish. With the number of the successively increasing
changes the difference in the magnitude of the change in the two stages
would always relatively diminish until it had quite vanished from our
perception; just in the same manner as we can distinguish a group of
three grains of corn from one composed of six, but not a heap of 103
grains from one containing 106 grains.
That the small systematic groups must have required a short period and
the large groups a long period of time for their formation requires no
special proof, but results immediately from the theory of descent.
All the foregoing considerations would, however, only hold good if
the transforming impulses were equal in strength, or, not to speak
figuratively, if the changes only occurred in equivalent portions of
the body, _i.e._ in such portions as those in which the changes are
of the same physiological and morphological importance to the whole
organism.
Now in the lower systematic groups this is always the case. Varieties,
species, and genera are always distinguished by only relatively small
differences; deep-seated distinctions do not here occur, as is implied
in the conception of these categories. The true cause of this is, I
believe, to be found in the circumstance that all changes take place
only by the smallest steps, so that greater differences can only arise
in the course of longer periods of time, within which a great number of
types (species) can, however, come into existence, and these would be
related by blood and in form in different degrees, and would therefore
form a systematic group of a higher rank.
Public-domain text, read in full here on John Shaqi.
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