Now if even under the most widely diverging conditions of life
adaptation of form is produced by relatively small, and to a
certain extent superficial, changes, we should expect less typical
transformations in the great majority of caterpillars which live
on the exterior of plants or in their softer parts (most of the
Micro-lepidoptera). The great diversity in the forms of caterpillars
depends essentially upon a different formation of the skin and its
underlying portions. The skin is sometimes naked, and can then acquire
the most diverse colours, either protective or conspicuous, or it may
develop offensive or defensive markings; in other cases it may be
covered with hairs which sting, or with spines which prick; certain
of its glands may develop to an enormous size, and acquire brilliant
colours and the power of emitting stinking secretions (the tentacles of
the _Papilionidæ_ and Cuspidate larvæ); by the development of warts,
angles, humps, &c., any species of caterpillar may be invested with the
most grotesque shape, the significance of which with respect to the
life of the insect is as yet in most cases by no means clear: _typical
portions_ are not, however, essentially influenced by these manifold
variations. At most only the form of the individual segments of the
body, and with these the shape of the whole insect, become changed
(onisciform larvæ of _Lycænidæ_), but a segment is never suppressed,
and even any considerable lengthening of the legs occurs but very
seldom (_Stauropus Fagi_).[194]
We may therefore fairly assert that the structure of larvæ is on the
whole remarkably uniform, in consequence of the uniformity in the
conditions of life. Notwithstanding the great variety of external
aspects, the general structure of caterpillars does not become
changed--it is only their outward garb which varies, sometimes in one
direction, and sometimes in another, and which, starting from inherited
characters, becomes adapted to the various special conditions of life
in the best possible manner.
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