Meanwhile, in the absence of further observations, we must admit that
the Paris Axolotls were not _Siredon Lichenoides_, but some nearly
allied and probably new species. But little information is furnished
by observing the course of the transformation, although it is at least
established that this Axolotl in its native habitat does not undergo
metamorphosis or does so as exceptionally as in Europe. Unfortunately
in his papers Duméril gives no precise statement respecting the
locality of this species imported from “Mexico”--it is probable that
he was himself unacquainted with it, so that I can only state on the
authority of Cope that Amblystoma has never been brought from south of
the provinces of Tamaulipas and Chilhuahua, _i.e._ south of the Tropic
of Cancer.[233]
This last statement, however, gives no certainty to the matter. Of much
more importance is the above-mentioned fact, that the true Axolotl
of the lakes about the city of Mexico does not, as a rule, become
transformed into an Amblystoma in that locality, although this species
in certain cases undergoes metamorphosis when in confinement. From
this circumstance and from the fact that the Paris Axolotl likewise
experienced but a very small percentage of metamorphosis in captivity,
we may venture to conclude that this species also, in its native
habitat, either does not become transformed at all or does so only
exceptionally.
But there is yet another body of facts which come prominently into
view on considering the history of the transformations. I refer to the
existence of numerous species of _Amblystoma_ in a natural state. In
the “Revision of the genera of _Salamandridæ_,” published some years
ago by Strauch,[234] this author, following Cope,[235] gives twenty
species of _Amblystoma_ as inhabiting North America. Although some
of these species are based on single examples, and consequently, as
Strauch justly remarks, “may well have to be reduced in the course
of time,” there must nevertheless always remain a large number of
species which live and propagate as true Amblystomas, and of which the
habitat extends from the latitude of New York to that of New Mexico.
There are therefore true species of _Siredon_ which regularly assume
the Amblystoma form under their natural conditions of life, and which
propagate in this form, while, on the other hand, there are at least
two species which, under their existing natural conditions of life,
always propagate as _Siredon_. It is but another mode of expression
for the same facts if we say that the Mexican Axolotl and the Paris
_Siredon_--whether this is _Lichenoides_ or some other species--stand
at a lower grade of phyletic development than those species of
_Amblystoma_ which propagate in the salamander form. No one can raise
any objection to this statement, while the alternative view maintained
by all authors contains a theory either expressed or implied which is,
as I believe, incorrect, viz. that the Mexican Axolotl _has remained_
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