at an inferior stage of phyletic development.
All zoologists[236] who have expressed an opinion upon the
transformation of the Axolotl, and who are not, like the first observer
of this fact, embarrassed by Cuvier’s views as to the immutability of
species, regard the phenomenon as though a species, which owing to some
special conditions had hitherto remained at a low stage of development,
had now through some other influences been compelled to advance to a
higher stage.
I believed for a long time that the phenomenon could not otherwise be
comprehended, so little was I then in a position to bring all the facts
into harmony with this view. Thus in the year 1872 I expressed myself
as follows[237]:--“Why should not a sudden change in all the conditions
of life (transference from Mexico to Paris) have a direct action on
the organization of the Axolotl, causing it suddenly to reach a higher
stage of development, such as many of its allies have already attained,
and which obviously lies in the nature of its organization--a stage
which it would perhaps itself have reached, although later, in its
native country? Or is it inconceivable that the sudden removal from
8000 feet above the sea (Mexican plateau) to the altitude of Paris, may
have given the respiratory organs an impetus in the direction of the
transformation imminent? In all probability we have here to do with the
direct action of changed conditions of life.”
That the substance of this last statement must still hold good is
obvious from the experiments previously described, which show that
by the application of definite external influences, we have it to
a certain extent in our power to produce the transformation. It is
precisely in this last point that there lies the new feature furnished
by these experiments.
But are we also compelled to interpret the phenomenon in the above
manner? _i.e._ as a sudden advance in the phyletic development of the
species occurring, so to speak, at one stroke? I believe not.
What first made this view appear to me erroneous, was the appearance
of the living Amblystomas bred from my Axolotl larvæ. These creatures
by no means differed from the Axolotls merely in single characters,
but they were distinct from the latter in their entire aspect; they
differed in some measure in all their parts, in some but slightly and
in other parts strongly--in brief, they had become quite different
animals. In accordance with this, their mode of life had become
completely modified; they no longer lived in the water, but remained
concealed by day among the damp moss of the vivarium, coming forth only
by night in search of food in dry places.
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