In the first place, the theory readily explains why the summer but not
the winter generations are capable of being transformed; the latter
cannot possibly revert to the _Prorsa_ form, because this is much the
younger. When, however, it happens that out of a hundred cases there
occurs one in which a chrysalis of the winter generation, having been
forced by warmth, undergoes transformation before the commencement of
winter, and emerges in the summer form,[18] this is not in the least
inexplicable. It cannot be atavism which determines the direction of
the development; but we see from such a case that the changes in the
first two generations have already produced a certain alteration in
the third, which manifests itself in single cases under favourable
conditions (the influence of warmth) by the assumption of the _Prorsa_
form; or, as it might be otherwise expressed, the _alternating_
heredity (of which we shall speak further), which implies the power
of assuming the _Prorsa_ form, remains latent as a rule in the winter
generation, but becomes _continuous_ in single individuals.
It is true that we have as yet no kind of insight into the nature of
heredity, and this at once shows the defectiveness of the foregoing
explanation; but we nevertheless know many of its external phenomena.
We know for certain that one of these consists in the fact that
peculiarities of the father do not appear in the son, but in the
grandson, or still further on, and that they may be thus transmitted
in a latent form. Let us imagine a character so transmitted that it
appears in the first, third, and fifth generations, remaining latent
in the intermediate ones; it would not be improbable, according to
previous experiences, that the peculiarity should exceptionally, i.e.,
from a cause unknown to us, appear in single individuals of the second
or fourth generation. But this completely agrees with those cases in
which “exceptional” individuals of the winter brood took the _Prorsa_
form, with the difference only that a cause (warmth) was here apparent
which occasioned the development of the latent characters, although we
are not in a position to say in what manner heat produces this action.
These exceptions to the rule are therefore no objection to the theory.
On the contrary, they give us a hint that after one _Prorsa_ generation
had been produced, the gradual interpolation of a second _Prorsa_
generation may have been facilitated by the existence of the first.
I do not doubt that even in the natural state single individuals of
_Prorsa_ sometimes emerge in September or October; and if our summer
were lengthened by only one or two months this might give rise to a
third summer brood (just as a second is now an accomplished fact),
under which circumstances they would not only emerge, but would also
have time for copulation and for depositing eggs, the larvæ from which
would have time to grow up.
Public-domain text, read in full here on John Shaqi.
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