The explanation of these facts is easily afforded by the foregoing
theory. According to this, both the winter forms must be regarded as
primary, and the _Marcellus_ form as secondary. But this last is not
yet so firmly established as _Prorsa_, in which reversion of the summer
generations to the _Levana_ form only occurs through special external
influences; whilst in the case of _Ajax_ some individuals are to be
found in every generation, the tendency of which to revert is still so
strong that even the greatest summer heat is unable to cause them to
diverge from their original inherited direction of development, or to
accelerate their emergence and compel them to assume the _Marcellus_
form. It is here beyond a doubt that it is not different external
influences, but internal causes only, which maintain the old hereditary
tendency, for all the larvæ and pupæ of many different broods were
simultaneously exposed to the same external influences. But, at the
same time, it is evident that these facts are not opposed to the
present theory; on the contrary, they confirm it, inasmuch as they
are readily explained on the basis of the theory, but can scarcely
otherwise be understood.
If it be asked what significance attaches to the duplication of the
winter form, it may be answered that the species was already dimorphic
at the time when it appeared in only one annual generation. Still,
this explanation may be objected to, since a dimorphism of this kind
is not at present known, though indeed some species exhibit a sexual
dimorphism,[20] in which one sex (as, for instance, the case of the
female _Papilio Turnus_) appears in two forms of colouring, but not a
dimorphism, as is here the case, displayed by both sexes.[21] Another
suggestion, therefore, may perhaps be offered.
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