Studies of Birds Killed in Nocturnal MigrationTordoff, Harrison Bruce
Science
Studies of Birds Killed in Nocturnal Migration
Tordoff, Harrison Bruce
Birds -- Migration
LINEAR MEASUREMENTS.--Taxonomists long have recognized in many species
that males differ in size from females. Less attention, until
recently, has been paid to the relative sizes of adult and immature
birds. Many taxonomists, however, seem to have had an uneasy suspicion
that immature birds are "untrustworthy" in comparison with adults, and
immatures have often been excluded from samples when recognizable.
Since, however, there are still relatively few reliably aged specimens
in collections, for the most part only those immature birds
immediately recognizable as such by obvious plumage differences (which
are often present only in juvenal plumage) have been excluded from
series. The majority of birds in first winter plumage so closely
resemble adults that the two ages have been included in series for
measurement. In most passerines these younger birds still bear the
juvenal feathers in wing and tail and are, in size of these important
parts, quite as "untrustworthy" as birds still in juvenal body
plumage. Even if a complete postjuvenal molt occurs we still should
not assume that first winter feathers are as long as adult winter
feathers without first determining that this is so. Although aware of
this problem, systematists until recently seemingly have been more or
less content to disregard it, or forced to do so for practical
reasons. Miller (1941:179) had little choice but to hope that size
differences between adult and immature juncos were unimportant. Behle
(1942:217) wrote of Horned Larks, _Eremophila alpestris_: "... the
plumages of first-year birds and adults seem indistinguishable, though
I have never quite satisfied myself that there are no differences in
lengths of rectrices and remiges." He added, with logic confusing to
us: "Since it is a difficult problem to determine the ages of horned
larks that have passed the postjuvenal molt, this similarity of
plumages is fortunate for the systematist."
In recent years, some workers have analyzed size differences between
adults and immatures. Sibley (1950:115) showed that adult Red-eyed
Towhees (_Pipilo erythrophthalmus_) had notably longer wings and tails
than immatures, and the same was demonstrated in Red Crossbills
(_Loxia curvirostra_) by Tordoff (1952). In work with jays
(_Aphelocoma_), Pitelka (1951:199) found that: "... in comparisons of
dimensions of sex and age groups within a given sample, although
magnitude of difference varies from one character to another, most of
the averages are successively smaller for first-year males and adult
and first-year females." He listed exceptions and concluded:
"Segregation [of sex and age classes] has proved to be of extreme
significance in an interpretation of individual and geographic
variation."
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