Subspeciation in the Kangaroo Rat, Dipodomys ordii: KU. Vol 1 No 23Setzer, Henry W.
Science
Subspeciation in the Kangaroo Rat, Dipodomys ordii: KU. Vol 1 No 23
Setzer, Henry W.
Kangaroo rats
Several heteromyids which have not been assigned to any genus are known.
Wilson (1939:36-37) recognized some from the Avawatz (Clarendonian) and
the Ricardo (Clarendonian). Another, possibly of the genus
_Diprionomys_?, from the Barstow (Barstovian) was described by Wood
(1935:197) as follows: "The general shape of the tooth as figured
strongly suggests either one of the most advanced species of _Dipodomys_
or else a Geomyid.... It is much more advanced than are any known
contemporary heteromyids, and compares fairly well with such late
Tertiary and Pleistocene geomyids as have been described. It certainly
is not referable to any known heteromyid genus other than _Dipodomys_,
and should probably be called a Geomyid." Wilson (_loc. cit._) refers to
these specimens as Dipodomyine (?) n. gen. and sp. If these specimens
referred to by Wood and Wilson are true heteromyids then a change in the
phylogenetic scheme proposed by Wood (1935) would be necessary. Wilson
(_loc. cit._) says, referring to the Avawatz specimen, "The cheek teeth
are very hypsodont but are apparently not persistent in growth,... Wide
enamel breaks are present in M/1 dividing the enamel into anterior and
posterior bands. The enamel of P/4 is complete in the present stage of
wear, but an examination of the tooth indicates that breaks would
develop with additional attrition at the buccal and lingual margins of
the metalophid, and at the buccal border of the protolophid. The incisor
is of the slender heteromyid type."
Wood (1935:118) in referring to the ancestry of _Cupidinimus_ with
regard to the grooving of the incisors says: "The philosophy of
evolution which would prohibit its derivation from _Mookomys_, because
of the grooved incisors in the latter genus, would require a separate
line leading back at least to the Lower Miocene."
In view of the above statements, it is conceivable that additional
material will be found carrying the dipodomyine line back into the
early Miocene. Perhaps the line involving _Mookomys_ and _Cupidinimus_
which was regarded by Wood as the line of descent, is merely an aberrant
side branch that parallels in its structures the main line of evolution
of the dipodomyines (Figure 1).
[Illustration: FIG. 1. Phylogeny of the Dipodomyines (modified after
Wood, 1935).]
As Wilson (1939:37) says: "Indeed it is hard to recognize such a form as
_Cupidinimus nebraskensis_ as directly ancestral to _Dipodomys_ in view
of the occurrence of the much more advanced Avawatz specimen in deposits
that are at most only slightly later than those in which the former is
found. The kangaroo rats were apparently much farther along in their
development by lower Pliocene time than heretofore supposed."
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