The racemes containing the flowers which were to be pollinated by hand
were covered with tarlatan before any of the flowers opened and were
kept covered except while being pollinated until the seeds were nearly
mature. This cloth has about twice as many meshes to the linear inch as
ordinary mosquito netting and served to exclude all insects that are
able to pollinate the flowers. When entire plants were to be protected
from all outside agencies, cages covered with cheesecloth, glass
frames, or wire netting were used.
A preliminary study of the pollination of _Melilotus alba_ and _M.
officinalis_ showed that both were visited by the same kinds of insects
and that both required the same methods of pollination in order to set
seed. On this account _M. alba_ was used in most of the experiments
reported in this bulletin. Where _M. officinalis_ was employed it is so
stated.
STRUCTURE OF THE FLOWERS OF MELILOTUS ALBA.
[Illustration: Fig. 1.--Different parts of the flower of
_Melilotus alba_: 1, Side view of the flower; 2, side view of the
flower with the carina and alæ slightly depressed; 3, side view of the
flower, showing the carina and alæ depressed sufficiently to expose
the staminal tube and the tenth free stamen; 4, ala; 5, ate and carina
spread apart to show their relative position and shape; 6, flower after
the petals have been removed, showing in detail the calyx and staminal
tube; 7, the staminal tube split open to show the relative size and
position of the pistil, _a_, Alæ; _b_, vexillum; _c_, carina; _d_,
calyx; _c_, stigma; _b_, anthers: _g_, tenth free stamen; _h_, digitate
process of the superior basal angle of an ala; _i_, depressions in the
ala; _j_, staminal tube; _k_, pistil.]
The racemes of _Melilotus alba_ contain from 10 to 120 flowers with
an average of approximately 50 per raceme for all of the racemes of a
plant growing under cultivation in a field containing a good stand.
The flower consists of a green, smooth, or slightly pubescent calyx
with 5-pointed lobes and with an irregular white corolla of five
petals. (Fig. 1.) The claws of the petals are not united nor are they
attached to the staminal tube which is formed by the union of the
filaments of the nine inferior stamens. As the claws of the alæ and
carina are not attacked to the staminal tube; the petals may be bent
downward sufficiently far so that many different kinds of insects may
secure without difficulty the nectar secreted around the base of the
ovary.
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