Produced by Chris Curnow, Joseph Cooper and the Online Distributed Proofreading Team at https://www.pgdp.net Taxonomy of the Chipmunks, Eutamias quadrivittatus and Eutamias umbrinus BY JOHN A. WHITE University of Kansas Publications Museum of Natural History Volume 5, No. 33, pp. 563-582, 6 figures in text December 1, 1953 University of Kansas LAWRENCE 1953 UNIVERSITY OF KANSAS PUBLICATIONS, MUSEUM OF NATURAL HISTORY Editors: E. Raymond Hall, Chairman, A. Byron Leonard, and Robert W. Wilson Volume 5, No. 33, pp. 563-582, 6 figures in text December 1, 1953 UNIVERSITY OF KANSAS Lawrence, Kansas PRINTED BY FERD VOILAND, JR., STATE PRINTER TOPEKA, KANSAS 1953 24-8966 Taxonomy of the Chipmunks, Eutamias quadrivittatus and Eutamias umbrinus By JOHN A. WHITE The differences in anatomy and color between many species of chipmunks are subtle, and refined techniques are required to discover them. When "measuring" chipmunks taxonomically, it is necessary to use a "chipmunk scale" and not, for example, a "pocket-gopher scale." In explanation, some species of pocket gophers closely allied to each other, and even some subspecies of the same species, differ markedly in color and in size and shape of parts of the skeleton; comparable differences are not so pronounced among many species of chipmunks. HISTORICAL SUMMARY Merriam (1905) was the first to show clearly that _Eutamias quadrivittatus_ is a distinct species, and pointed out that _E. amoenus operarius_ (= _E. minimus operarius_) is a small species which resembles, and is found in some areas together with, _E. quadrivittatus_. Howell (1929) placed under _E. quadrivittatus_ the following subspecies: _E. q. quadrivittatus_, _E. q. hopiensis_, _E. q. inyoensis_, _E. q. frater_, _E. q. sequoiensis_, and _E. q. speciosus_. Hardy (1945) placed _E. adsitus_ under _E. quadrivittatus_ as _E. q. adsitus_, and Kelson (1951) placed _E. umbrinus_ under _E. quadrivittatus_ as _E. q. umbrinus_. Johnson (1943) re-established _E. speciosus_ as a separate species, and in California left only _E. q. inyoensis_ in _E. quadrivittatus_. Thus, since 1943 the recognized subspecies of _E. quadrivittatus_ have been: _E. q. quadrivittatus_, _E. q. hopiensis_, _E. q. inyoensis_, _E. q. nevadensis_, _E. q. umbrinus_, and _E. q. adsitus_.