If, then, every male is driven by inherited impulse to seek the
appropriate breeding ground each recurring season; if, having arrived
there, it is driven to seek a position of its own; if, in order to
secure isolation it is obliged to attack other males or to ward off the
attacks of intruders; if, in short, success can only be attained
providing that the inherited nature is so adjusted that the bird can
accomplish all that is here demanded--what will be the general result?
That the individual will rear its offspring in safety and that they will
inherit the peculiarities of their parents, enabling them, in their
turn, to procreate their kind; all this will certainly follow. We are
not concerned, however, at the moment, with the direct effect upon the
individual, but with the consequences that will accrue to the species as
a whole.
Now certain facts are presented to observation which enable us not only
to understand the nature of the change that is wrought in the history of
the species, but to foreshadow, with no small degree of certainty, the
extent of that change. I suppose that it has come within the experience
of most of us to observe, at one time or another, the ebb and flow of a
given species in a given district. Some favourite haunt is deserted for
a year, or for a term of years, and is then revisited; or, if it is
always occupied, the number of inhabitants fluctuates--plenty of pairs
in this season, only a few in that. Many intricate relationships, both
external and internal, contribute towards this state of affairs.
Fluctuation in a downward direction, or temporary extinction, is brought
about by changes in the physical world, by changes in the available
supply of food, by the increase of enemies, or by adverse climatic
conditions; whilst fluctuation in an upward direction, though due
indirectly to a combination of circumstances in the external world
favourable to the survival of large numbers of individuals, is directly
determined by the impulse to seek isolation. As individuals of
different species establish themselves, and form kingdoms and lesser
kingdoms, we can watch the gradual quickening into life of moorland and
forest and we can observe the manner in which it all comes to pass.
Males that for weeks or months have lived in society, drifting from
locality to locality according to the abundance of food or its scarcity,
now set forth alone and settle first here and then there in search of
isolation. Lapwings settle in the water meadows, and, finding themselves
forestalled, pass on in search of other ground; Blackbirds arrive in a
coppice or in a hedgerow and, meeting with opposition, disappear; and
the Curlew, wandering with no fixed abode but apparently with a fixity
of purpose, searches out the moorland where it can find the particular
environmental conditions to which its inherited nature will respond. In
fact, wherever we choose to look, we can observe in a general way the
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