gradual appropriation of breeding ground; and if we fix our attention
upon particular males, we can watch the method by which success or
failure is achieved.
On more than one occasion I have watched the efforts of Reed-Buntings to
appropriate territories in a marsh that was already inhabited. Sometimes
their efforts met with success, at other times with failure. In the
former case, the males, whose ground was intruded upon, were severally
forced to yield part of their holding and were thus left in possession
of a smaller area. The success of the intruder seemed to depend upon
persistent determination, rather than upon superior skill in battle.
Recently I had an opportunity of observing the intrusion of a male
Willow-Warbler upon ground already occupied. By persistent effort it
succeeded in appropriating one half of the territory of its rival. The
intruder occupied some trees on the outskirts of the territory it was
invading, and used them as a base from which it made repeated efforts to
enter the ground of its rival. These efforts were time after time
frustrated. No sooner did it leave its base than it was seen and
intercepted, or else attacked; and no matter from which direction it
attempted to effect an entrance, its efforts, for a time, were all to no
purpose. The fighting was of a determined character, and after each
attack the owner of the territory showed signs of great excitement, and,
sitting upright upon a branch, spread and waved its wings, which is the
specific emotional manifestation during the period of sexual activity.
Eventually the intruding male succeeded by persistent effort in
appropriating part of the occupied ground.
Thus we can actually witness the efforts of the individual to isolate
itself from members of its own kind, and can observe the immediate
consequences that follow from success or from failure. And from these
consequences we can infer that, within a certain range but in accordance
with the relative abundance of the species that dwell in it, every
corner of the available breeding ground will be explored and every
situation that evokes the appropriate response will be occupied.
Moreover, since the annual dispersion is not merely a repetition in this
season of that which occurred in a previous one, a progressive increase
in the area occupied will follow. Yet, if the majority of species desert
their breeding ground so soon as reproduction is ended, how can this be?
An answer to the question will be found in the fact that a bird has an
innate capacity to return to the neighbourhood of its birthplace, or to
the place wherein it had previously reared offspring--which means that
the results of prior process persist as the basis and starting-point of
subsequent process.
Public-domain text, read in full here on John Shaqi.
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