The disadvantages which the late arrivals have to face are therefore
great, and it is probable that the percentage which attain to
reproduction will on the average be somewhat lower than the percentage
in the case of the earlier arrivals. The district in which my
observations have been made lies well within the limits of the breeding
range of most of our common species, and it is not surprising that I
should have met with little evidence of failure to breed as a result of
failure to secure territory. Some interesting information was supplied
to me, however, by the late Robert Service. He found, in certain seasons
in Dumfriesshire, flocks of from ten to fifty unmated Sedge-Warblers,
which, from the time of their arrival in May until the middle of July,
haunted reed-filled spaces along stagnant streams. These flocks appeared
to him to be composed of loosely-attached individuals of a migrant flock
that had failed to find things congenial enough to entice them to
disperse. But may they not have been composed of males that had failed
to secure territories, or of females that had failed to discover males
in possession of territories, or of both?
We have seen that, in the case of many species, each male establishes
itself in a particular place at the commencement of the breeding season,
even though this may mean a partial or perhaps a complete severance from
former companions. We must now discuss this fact in greater detail
because it is opposed to the views often held regarding the sexual
behaviour of birds, and is manifestly of importance when considering
the theory of breeding territory.
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