Lapwings, as is well known, collect in flocks during the winter months,
and these flocks, which sometimes reach vast proportions, are to be
found on tidal estuaries, water meadows, arable land, and such like
places, according to the prevailing climatic conditions. This flocking
may contribute towards survival, and may therefore be the result of
congenital dispositions which have been determined on biological
grounds. On the other hand, since food at that season is only to be
obtained in a limited number of situations, the birds may be simply
drawn together by accident. In the former case the behaviour would be
instinctive, in the latter, though accidental at first, recurrent
repetition would tend to make it habitual; but in either case the
impulse to accompany the flock must be a powerful one, for on the one
hand it would depend upon inherited, and on the other hand upon
acquired, connections in the nervous system. Now observe that soon
after the flock arrived in the meadow, single males detached themselves;
there was no hesitation, they just retired from their companions and
settled in their respective territories. They were not expelled, for if
their leaving had been compulsory much commotion would have preceded
their departure, and their return would certainly not have been
welcomed. A reference to the plan will make the position clearer; the
neutral zone inhabited by the flock is there shown as situated in one
corner of the meadow, the territories that fell under observation are
plotted out as far as possible to scale, and the more important zones of
conflict are also marked.
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