Which of these three shall we choose? The first by itself requires but
little consideration; for though it might explain the initial visit, it
cannot account for the persistency with which the plot of ground is
afterwards resorted to. Supposing, however, that we combine the first
and the second; supposing, that is to say, we assume, for the purpose of
argument, that the initial visit is fortuitous, and that constancy is
supplied by habit formation--would that be a satisfactory
interpretation? It is a simple one, inasmuch as it only requires that a
male shall alight by chance in a particular place for a few mornings in
succession in order that the process may be set in motion. Now an
essential condition of habit formation is recurrent repetition; given
this repetition and, it is true, any mode of activity is liable to
become firmly established. But how can we explain the repetition? Even
if we are justified in assuming that the initial visit is purely an
accidental occurrence, we cannot presume too far upon the laws of chance
and assume that the repetition, at first, is also fortuitous.
So that we come back to the congenital basis, the last of our three
propositions. And it will, I think, be admitted that the facts give us
some grounds for believing that the securing of the territory has its
root in the inherited constitution of the bird. In comparing the
behaviour of the migratory male with that of the resident, attention was
drawn to the manner in which the occupation of a territory was effected:
the former bird, it may be remembered, established itself without delay,
whereas the latter did so only by degrees, and the difference was
attributed to the incidence of migration which required a closer
correspondence between organic process and external environment. But the
significance for us just now lies in the fact that the definiteness,
which accompanies the initial behaviour of the migratory male in
relation to the territory, cannot have been acquired by repetition; for
this reason, that when the male occupies its space of ground at the end
of its long and arduous journey, it does so without preparation or
experiment, even without hesitation, as if aware that it was making good
the first step in the process of reproduction. No doubt, if it happened
to be an individual that had already experienced the enjoyment of
reproduction, it might be aware of the immediate results to be achieved
and act accordingly. But among the hosts of migrants that one observes,
there must be many males which have not previously mated; and yet, upon
arrival, they all behave in a similarly definite manner--so that
experience cannot well be the primary factor in the situation. If, then,
the essential condition of habit formation is absent and experience is
eliminated, there is nothing left but racial preparation to fall back
upon.
Public-domain text, read in full here on John Shaqi.
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