In attributing the rivalry to the presence of the female, it is assumed
that males are in a preponderance, and that consequently two or more are
always ready to compete for a mate. Her presence is presumably the
condition under which his pugnacious nature is rendered susceptible to
its appropriate stimulus, the stimulus being, of course, supplied by the
opponent. There would be nothing against this interpretation if it were
in accord with the facts; but it can, I think, be shown that the males
are just as pugnacious and the conflicts just as severe even when the
question of securing a mate is definitely excluded; and I shall now give
the evidence which has led me to this conclusion.
In the previous chapter we had occasion to refer to the difference in
the times of arrival of the male and female migrants, and we came to the
conclusion, it may be remembered, that this was a fact of some
importance, because it gave us a clue to the meaning of much that was
otherwise obscure in their behaviour. But it is also of importance in
connection with the particular aspect of the problem which we now have
in view, for if it can be shown that males, when they first reach their
breeding grounds, are even then intolerant of one another's presence, if
their actions and attitudes betray similar symptoms of quasi-conation,
if disputes are rife and the struggles of a kind to preclude all doubt
as to their reality, then it is manifest that in such cases their
intolerance cannot be due to the presence of the female.
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