But even granting that this disposition forms part of the hereditary
equipment of the bird, how is the process of reproduction furthered? The
mere fact of remaining in or about a particular spot cannot render the
attainment of reproduction any less arduous, and may indeed add to the
difficulties, for any number of individuals might congregate together
and mutually affect one another's interests. A second disposition comes,
however, into functional activity at much the same stage of sexual
development, and manifests itself in the male's intolerance of other
individuals. And the two combined open up an avenue through which the
individual can approach the goal of reproduction. In terms of the theory
I shall refer to this second disposition as the one which is concerned
with the defence of the territory.
Broadly speaking, these two dispositions may be regarded as the basis
upon which the breeding territory is founded. Yet inasmuch as the
survival value of the dispositions themselves must have depended upon
the success of the process as a whole, it is manifest that peculiar
significance must not be attached to just the area occupied, which
happens to be so susceptible of observation; other contributory factors
must also receive attention, for the process is but an order of
relationships in which the various units have each had their share in
determining the nature and course of subsequent process, so that, as Dr
Stout says, when they were modified, it was modified.
Now the male inherits a disposition which leads it to remain in a
restricted area, but the disposition cannot determine the extent of that
area. How then are the boundaries fixed? That they are sometimes adhered
to with remarkable precision, that they can only be encroached upon at
the risk of a conflict--all of this can be observed with little
difficulty. But if we regard them as so many lines definitely delimiting
an area of which the bird is cognisant, we place the whole behaviour on
a different level of mental development, and incidentally alter the
complexion of the whole process. It would be a mistake, I think, to do
this. Though conscious intention as a factor may enter the situation,
there is no necessity for it to do so; there is no necessity, that is to
say, for the bird to form a mental image of the area to be occupied and
shape its course accordingly. The same result can be obtained without
our having recourse to so complex a principle of explanation, and that
by the law of habit formation. In common with other animals, birds are
subject to this law in a marked degree. An acquired mode of activity
becomes by repetition ingrained in the life of the individual, so that
an action performed to-day is liable to be repeated to-morrow so long as
it does not prejudice the existence or annul the fertility of the
individual.
Public-domain text, read in full here on John Shaqi.
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