When the food has been distributed through the intestine so as to
present the appearance shown in Figure 1, a noticeable feature in most
or all of the loops is the total absence of movement. If the animal
remains quiet, however, only a few moments elapse before peculiar
motions appear in one or another of the loops, or perhaps in several,
and last for some time. These motions consist in a sudden division
of one of the long, narrow masses of food into many little segments
of nearly equal size; then these segments are again suddenly divided
and the neighbouring halves unite to make new segments, and so on, in
a manner to be more fully described. I have called this process the
rhythmic segmentation of the intestinal contents. Further observation
reveals peristalsis here and there, and under certain circumstances the
typical swaying movements may be seen. All these phenomena are now to
be considered in detail.
[Illustration:
FIGURE 1.--Appearance of food in the intestines 5¾ hours after eating.
This and other radiographs reduced two-thirds.]
[Illustration:
FIGURE 2.--Diagram representing the process of rhythmic segmentation.
Lines 1, 2, 3, 4 indicate the sequence of appearances in the loop.
The dotted lines mark the regions of division. The arrows show the
relation of the particles to the segments they subsequently form.]
_Rhythmic segmentation of the intestinal contents._--This is by far the
most common and the most interesting mechanical process to be seen in
the small intestine. The nature of the process may best be understood
by referring to the diagram in Figure 2. A string-like mass of food
is seen lying quietly in one of the intestinal loops (line 1, Fig.
2). Suddenly an undefined activity appears in the mass, and a moment
later constrictions at regular intervals along its length cut it into
little ovoid pieces. The solid string is thus quickly transformed,
by a simultaneous sectioning, into a series of uniform segments. A
moment later each of these segments is divided into two particles,
and immediately after the division neighbouring particles (as _a_
and _b_, line 2, Fig. 2) rush together, often with the rapidity of
flying shuttles, and merge to form new segments (as _c_, line 3, Fig.
2). The next moment these new segments are divided, and neighbouring
particles unite to make a third series, and so on. At the time of the
second segmentation (line 3, Fig. 2) the end particles are left small.
Observation shows that these small pieces are not redivided. The end
piece at A simply varies in size with each division; at one moment it
is left small, at the next moment it is full size from the addition
of a part of the nearest segment, and a moment later is the small bit
left after another division. The end piece at B (probably the rear
of the mass) shoots away when the end mass is divided, and is swept
back at each reunion to make the large end mass again, only to be
shot away and swept onward with each recurrence of the constrictions.
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