Thus the process of repeated segmentation continues, with the little
particles flitting towards each other and the larger segments shifting
to and fro, commonly for more than half an hour without cessation.
From the beginning to the end of a period of segmentation the food
is seen to have changed its position in the abdomen to only a slight
extent; whether this change is a passing of the food along the loop,
or a movement of the loop itself, it is impossible to tell from the
shadows on the screen. The change of position, however, is much less
conspicuous than the lively division and redivision which the mass
suffers so many times from the busy, shifting constrictions.
From this typical form of rhythmic segmentation there are several
variations. Sometimes, and especially when the mass of food is thick,
the constrictions do not make complete divisions and are so far apart
that the intermediate portions are relatively large. Moreover, the
constrictions do not take place in the middle of each portion, but
near one end; thus each portion is constricted, not into halves,
but into thirds. If a little pointer is placed at the middle of a
segment, when the segments are completely divided into halves, in a few
seconds the pointer will be in the middle of the clear space between
two segments; but in a few seconds more the first phase will return
and the pointer will again be indicating a segment,--two operations
intervene between similar phases. When, however, the portions are
constricted into thirds, the indicator shows it, since three operations
intervene between similar phases. The manner of these changes is made
clearer by reference to the diagram in Figure 3. That each portion is
constricted into three pieces is proved also by watching the gradual
reduction of the portion at the left end of line 1 through lines 2
and 3, and also in the gradual formation of a full-sized portion at
the right end of lines 2, 3, and 4. When food undergoing this process
is watched, it appears to be affected by a series of constrictions,
each of which starts at one end of the mass and marches through to the
other end, leaving its impress at short intervals along the length. The
progression of the dotted lines from right to left in _a_, _b_, _c_,
and _d_, etc., Fig. 3, gives a notion of these advancing constrictions.
Another variation of the segmentation is shown in Figure 4. In this
type there are evidently divisions and subdivisions, _i.e._, one more
operation between the appearance and the reappearance of the same phase
than is present in the simple division of the small segments in a long
string of food (Fig. 2). This form of segmentation is fairly typical
for the constrictions seen in food advancing through the intestine.
Sometimes the divisions occur in the middle of a long string of food
and leave the ends wholly unaffected.
Public-domain text, read in full here on John Shaqi.
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