With the exception of the first two, to be discussed later, these are all essentially alike and the fourth will be described as a type (Fig. 17). Its centrum is cylindrical or somewhat hourglass shaped, concave anteriorly and convex posteriorly; it is not completely fused with the neural arch but is united with it by sutures. From the anterior end of the ventral surface of the centrum projects downward and forward a small plow-shaped process, the hypapophysis. On each side of the centrum, near its anterior end, is a facet with which the lower branch (capitulum) of the rib articulates. The neural arch is strongly developed and is extended dorsally into a prominent neural spine and on each side as a short, blunt, transverse process with which the tubercle or upper branch of the rib articulates. Posteriorly the arch is notched on each side to form the openings for the exit of the spinal nerves. Projecting dorsally and anteriorly from the arch are two short processes which bear the medially and dorsally facing prezygapophyses (Fig. 17). Just caudad to these processes are somewhat shorter processes that bear the laterally and ventrally facing postzygapophyses (Fig. 17, 6). The _atlas_, as in other vertebrates, is highly specialized. It consists (Fig. 17), even in the adult animal, six feet or more in length, of four distinct portions, a ventral (4), a dorsal (1), and two lateral (2) parts. The ventral portion is relatively more massive than in most animals; its anterior surface is concave and forms the main part of the articular surface for the occipital condyle of the skull. Its postero-dorsal surface articulates with the odontoid process of the axis. On its postero-lateral surfaces are the facets for articulation with the first ribs, which, unlike the other cervical ribs, have but one articular surface. Articulating dorsally with this ventral element of the atlas are the two rather heavy lateral elements which form the neural arch. Anteriorly they form the lateral parts of the articular surface for the condyle and dorsally they unite for a short distance with each other. Projecting ventrally from the posterior part of their dorsal portion are the small postzygapophyses. Ventrally and laterally they articulate with the odontoid process (Fig. 17, 3). Projecting dorsad and cephalad from the dorsal surface of these lateral elements is the dorsal element of the atlas (Fig. 17, 1), the pro-atlas, which may not be properly a part of the vertebral column at all, since it is said to be merely a membrane bone. Gadow says it is the detached neural spine of the atlas. It is thin and triangular in shape, resembling in contour a large, mammalian epiglottis. It forms an arch over the space between the skull and the front of the atlas proper. Reynolds calls it the pro-atlas.