The alligator and its alliesReese, A. M. (Albert Moore)
Science
The alligator and its allies
Reese, A. M. (Albert Moore)
Alligators
_The Lateral Aspect_ (Fig. 21). As will be seen by the figure,
practically all of the bones visible in this view have already been
described, except those of the mandible, which will be described
separately. At the base of the skull are, however, two bones, the basi-
and alisphenoid, that have not been described and that show as well
in this as in any other view. The _basisphenoid_ (just below v and
hidden by the pterygoid) was mentioned in connection with the quadrate,
with whose posterior margin it articulates. It is an unpaired bone
of very irregular shape. Anteriorly it is flattened out to form the
rostrum, a rectangular process that forms the posterior part of the
interorbital septum; in fact it is the only part of the septum present
in a prepared skull, since the rest is cartilaginous. Dorso-laterally
the basisphenoid articulates with the alisphenoid; posteriorly with
the basioccipital; ventrally with the pterygoid; and posteriorly with
the exoccipital and basioccipital. On the dorsal surface of the
basisphenoid is the pituitary fossa, not seen, of course, in this view
of the skull.
The _alisphenoids_ (crossed by the dotted line from V) are a pair of
very irregular bones that form most of the antero-lateral walls of
the brain case. They articulate dorsally with the parietal, frontal,
and postfrontal; ventrally with the basisphenoid and pterygoid; and
posteriorly with the quadrate and some of the otic bones not visible in
this view. Between it and the quadrate, plainly visible in this view,
is a large opening, the _foramen ovale_ (V), through which, according
to Reynolds, the trigeminal nerve passes. In the middle line, directly
under the frontal bone, is an opening between the anterior wings of
the two alisphenoids, for the exit of the optic nerves. Ventrad and
caudad to this opening, and sometimes continuous with it, is another
large foramen, just dorsad to the rostrum, for the exit, according to
Reynolds, of the oculomotor and abducens nerves. Projecting caudad
is seen the rounded condylar part of the _basi-occipital_ (14) to
be described later, and dorso-cephalad to this is a part of the
exoccipital (13) in which four foramina may be seen; of the dorsal
three the one nearest the condyle and foramen magnum is for the exit
(Reynolds) of the hypoglossal nerve (XII); slightly dorso-cephalad to
this is one for the vagus nerve (X); between these two is a very small
one for a vein; the largest and ventrally located foramen is for the
entrance of the internal carotid (15). Another large foramen in the
exoccipital bone will be seen and described in connection with the
posterior view of the skull. Dorsal to the quadrate and largely bounded
by it is the wide _external auditory meatus_ (16), which leads into the
tympanic cavity. This cavity is complicated by a number of canals that
lead from it in various directions. Overhanging the cavity and meatus
is the squamosal bone, described in connection with the dorsal aspect
of the skull.
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