=The Metatarsus.= _Location._—The metatarsal bone extends downward and forward. In birds it consists of one bone, which articulates superiorly with the inferior extremity of the tibia. On the distal end it has a threefold trochlear arrangement which articulates with the three principal digits. _Description._—The proximal extremity posteriorly has a process which may be considered as a consolidation of originally separate metatarsal bones. The process at the supero-posterior part of the metatarsus is called the _hypotarsus_ of the _tarso-metatarsus_, through which extends a canal called the _hypotarsal canal_, and which gives passage to flexor tendons. The proximal end of the metatarsus is ossified from one center of ossification forming an epiphysis which caps the ends of the three original metatarsal bones that coalesce, first with one another, then with the epiphysis, thus forming a single compound bone. Above and just to the inside of the metatarsus there may occur a small bone which is imbedded in ligaments and articulates with the inner proximal surface of the metatarsus. This has been called by Gadow a _sesamoid bone_. The shaft shows _tendonal grooves_ which are best marked on the posterior surface. The shaft is rounded at the sides and flattened on the anterior and posterior surfaces. At the juncture of the middle and inferior thirds of the metatarsus there is a conical process turned slightly backward, which serves as a base for the _spur_. The spur is a horny structure. At the distal extremity occurs the threefold trochlear arrangement mentioned above, which incloses the fossa intercondyloidea. The inner trochlea is the broadest, and the outer the narrowest. The inner trochlea articulates with the proximal end of the second, the middle with the third, and the outer with the fourth toe. =The Phalanges.= _Location._—Most domestic fowls are provided with four digits, or toes; the Houdan and Dorking are provided with five. In fowls with four toes the three principal toes, the second, the third, and the fourth, are directed forward, and the first, or hallux, is directed backward. _Description._—The last phalanx of each toe, called the ungual phalanx, is slightly curved downward, is pointed anteriorly, like the claw of the cat or dog, and is covered with a horny sheath. The articular ends of the joints of the phalanges are enlarged. The base of the basal phalanx has two enlargements, the superior-inferior tubercles, or tuberculum superius et inferius, between which is located the fossa articularis transversa. Laterally the head has two condyles, condyli laterales, which are divided by the sulcus longitudinalis. The bodies of the phalanges vary in form; they are superiorly rounded, but inferiorly rounded, flat, or even somewhat concave. The distal extremity has an articular trochlea.