Lastly, the Lower Silurian Rocks have yielded a vast number of chambered shells, referable to animals which belong to the same great division as the Cuttle-fishes (the _Cephalopoda_), and of which the Pearly Nautilus is the only living representative at the present day. In this group of _Cephalopods_ the animal possesses a well-developed external shell, which is divided into chambers by shelly partitions ("septa"). The animal lives in the last-formed and largest chamber of the shell, to which it is organically connected by muscular attachments. The head is furnished with long muscular processes or "arms," and can be protruded from the mouth of the shell at will, or again withdrawn within it. We learn, also, from the Pearly Nautilus, that these animals must have possessed two pairs of breathing organs or "gills;" hence all these forms are grouped together under the name of the "Tetrabranchiate" Cephalopods (Gr. _tetra_, four; _bragchia_, gill). On the other hand, the ordinary Cuttle-fishes and Calamaries either possess an internal skeleton, or if they have an external shell, it is not chambered; their "arms" are furnished with powerful organs of adhesion in the form of suckers; and they possess only a single pair of gills. For this last reason they are termed the "Dibranchiate" Cephalopods (Gr. _dis_, twice; _bragchia_, gill). No trace of the true Cuttle-fishes has yet been found in Lower Silurian deposits; but the Tetrabranchiate group is represented by a great number of forms, sometimes of great size. The principal Lower Silurian genus is the well-known and widely-distributed _Orthoceras_ (fig. 55). The shell in this genus agrees with that of the existing _Pearly Nautilus_, in consisting of numerous chambers separated by shelly partitions (or septa), the latter being perforated by a tube which runs the whole length of the shell after the last chamber, and is known as the "siphuncle" (fig. 56, s). The last chamber formed is the largest, and in it the animal lives. The chambers behind this are apparently filled with some gas secreted by the animal itself; and these are supposed to act as a kind of float, enabling the creature to move with ease under the weight of its shell. The various air-chambers, though the siphuncle passes through them, have no direct connection with one another; and it is believed that the animal has the power of slightly altering its specific gravity, and thus of rising or sinking in the water by driving additional fluid into the siphuncle or partially emptying it. The _Orthoceras_ further agrees with the Pearly Nautilus in the fact that the partitions or septa separating the different air-chambers are simple and smooth, concave in front and convex behind, and devoid of the elaborate lobation which they exhibit in the Ammonites; whilst the siphuncle pierces the septa either in the centre or near it. In the Nautilus, however, the shell is coiled into a flat spiral; whereas in _Orthoceras_ the shell is