The Appendages, Anatomy, and Relationships of TrilobitesRaymond, Percy E. (Percy Edward)
Science
The Appendages, Anatomy, and Relationships of Trilobites
Raymond, Percy E. (Percy Edward)
Trilobites
_Possible preservations of extensors and flexors in Ceraurus_.--Among
Doctor Walcott's sections are four slices which I should not like to
use in proving the presence of longitudinal muscles, but which may be
admitted as corroborative evidence. Two of these transverse sections
(Nos. 114 and 199) show a dorsal and a ventral pair of dark spots in
positions which suggest that they represent the location of the dorsal
and ventral muscles, while two others (Nos. 131 and 140) show only the
upper pair of spots. The chief objection to this interpretation is
that it is difficult to imagine how the muscles could be so replaced
that they happen to show in the section. Both the sections showing all
four spots are evidently from the anterior part of the thorax, as they
show traces of the abdominal sheath, which seems to be squeezed
against the inside of the axial lobe, with the muscles (?) forced out
to the sides. The ventral pair lie just inside the appendifers, but
even if they are sections of muscles, all four are probably somewhat
out of place.
_Hypostomial Muscles._
The hypostoma fits tightly against the epistoma, or the doublure when
the epistoma is absent, but in no trilobite has it ever been seen
ankylosed to the dorsal test, and its rather frail connection
therewith is evidenced by the relative rarity of specimens found with
it in position. That the hypostoma was movable seems very probable,
and that it was held in place by muscles, certain. The maculæ were
always believed to be muscle scars until Lindstroem (1901, p. 8)
announced the discovery by Liljevall of small granules on those of
_Goldius polyactin_ (Angelin). These were interpreted as lenses
of eyes by Lindstroem, who tried to show that the maculæ of all
trilobites were functional or degenerate eyes. Most palæontologists
have not accepted this explanation, and since the so-called eyes cover
only a part of the surface of the maculæ, it is still possible to
consider the latter as chiefly muscle-scars.
In Lindstroem's summary (1901, pp. 71, 72) it is admitted that the
globular lenses are found only in _Bronteus_ (_Goldius_) (three
Swedish species only) and _Cheirurus spinulosus_ Nieszkowski, while
the prismatic structure supposed to represent degenerate eyes was
found in eleven genera (Asaphidæ, Illænidæ, Lichadidæ). All of these
are forms with well developed eyes, and Lindstroem himself points out
that the appearance of actual lenses in the hypostoma was a late
development, long after the necessity for them would appear to have
passed.
The use of the hypostoma has been discussed by Bernard (1892, p. 240)
and extracts from his remarks are quoted:
Public-domain text, read in full here on John Shaqi.
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