The Appendages, Anatomy, and Relationships of TrilobitesRaymond, Percy E. (Percy Edward)
Science
The Appendages, Anatomy, and Relationships of Trilobites
Raymond, Percy E. (Percy Edward)
Trilobites
The kinds of sediments in which trilobites are entombed have so far
afforded little evidence as to their habitat. Frech (Lethæa
palæozoica, 1897-1902, p. 67 _et seq._) who has collected such
evidence as is available on this subject, places as deeper water
Ordovician deposits the "Trinucleus-Schiefer" of the upper Ordovician
of northern Europe and Bohemia, the "Triarthrus-Schiefer" of America,
the "Asaphus-Schiefer" of Scandinavia, Bohemia, Portugal, and France,
and the Dalmania quartzite of Bohemia. .
_Cryptolithus_ and _Triarthrus_, although not confined to such
deposits, are apt to occur chiefly in very fine-grained shales, in
company with graptolites. These latter are distributed by currents
over great distances within short periods. It is somewhat curious that
the nearly blind burrowing Trinucleidæ, the dysphotic, large-eyed
Remopleuridæ and Telephus, the blind nektonic Agnostidæ and Dionide,
and the planktonic graptolites should go together and make up almost
the entire fauna of certain formations. Yet, when the life history of
each type is studied, a logical explanation is readily at hand, for
all have free-swimming larvæ.
A list of the methods of life noted above is given by way of summary,
with examples.
{Planktonic {Primarily Earliest protaspis of all trilobites
{ {Secondarily _Deiphon_, _Odontopleura_, etc.
{
Pelagic { {Primarily Later protaspis of all trilobites.
{ { _Naraoia_
{ {
{ { {Probably many thin-shelled
{ { { trilobites with large pygidia
{ { { (only partially nektonic)
{Nektonic {Secondarily {Cyclopygidæ }
{Remopleuridæ } (nektonic dysphotic)
{Crawlers and
{ slow swimmers Most trilobites with small pygidia.
{ _Triarthrus_, _Paradoxides_, etc.
Benthonic {Crawlers and Most trilobites with large pygidia.
{ active swimmers _Isotelus_, _Dalmanites_, etc.
{
{Crawlers, slow
{ swimmers, and Trinucleidæ, Harpedidæ,
{ burrowers some Mesonacidæ, etc.
FOOD AND FEEDING METHODS.
This subject has been less discussed than the methods of locomotion.
The study of the appendages has shown that while the mouth parts were
not especially powerful, they were at least numerous, and sufficiently
armed with spines to shred up such animal and vegetable substances as
they were liable to encounter. It having been ascertained that the
shape of the glabella and axial lobe furnishes an indication of the
degree of development of the alimentary canal it is possible to infer
something of the kind of food used by various trilobites.
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