The Branchiopoda were very highly specialized as early as Middle Cambrian time, the carapace of the Notostraca being fully developed and the abdomen limbless. Some (_Burgessia_) had numerous segments, but most had relatively few. The most striking point about them, however, is that so far as is known none of them had phyllopodan limbs. While the preservation is in most cases unsatisfactory, such limbs as are preserved are trilobite-like, and in the case of _Burgessia_ there can be no possible doubt of the structure. Another interesting feature is the retention by _Yohoia_ of vestiges of pleural lobes. The Middle Cambrian Branchiopoda are more closely allied to the Trilobita than are the modern ones, but still the subclass is not so closely related to that group as has been thought. Modern _Apus_ is certainly much less like a trilobite than has been supposed, and very far from being primitive. The Branchiopoda of the Middle Cambrian could have been derived from the trilobites by the loss of the pleural lobes, the development of the posterior margin of the cephalon to form a carapace, and the loss of the appendages from the abdominal segments. Modern branchiopods can be derived from those of the Middle Cambrian by the modification of the appendages through the reduction of the endopodite and exopodite and the growth of the endites and exites from the proximal segments. Carpenter (1903, p. 334), from his study of recent crustaceans, has already come to the conclusion that the Branchiopoda are not the most primitive subclass, and this opinion is strengthened by evidence derived from the Trilobita and from the Branchiopoda of the Middle Cambrian. COPEPODA. The non-parasitic Eucopepoda are in many ways much nearer to the trilobites than any other Crustacea. These little animals lack the carapace, and the body is short, with typically ten free segments and a telson bearing caudal furcæ. The head is composed of five segments (if the first thoracic segment is really the fused first and second), is often flattened, and lacks compound eyes. Pleural lobes are well developed, but instead of being flattened as in the trilobite, they are turned down at the sides or even incurved. A labrum is present. The antennules, antennæ, and mandibles are quite like those of trilobites. The antennules are very long and made up of numerous segments. The antennæ are biramous, the junction between the coxopodite and basipodite is well marked, and the endopodite consists of only two segments. The mandibles are said to "retain more completely than in any other Crustacea the form of biramous swimming limbs which they possess in the nauplius." The coxopodites form jaws, while both the reduced endopodite and exopodite are furnished with long setæ. The maxillulæ are also biramous, but very different in form from those of the trilobite, and the maxillæ are phyllopodan.