The Appendages, Anatomy, and Relationships of TrilobitesRaymond, Percy E. (Percy Edward)
Science
The Appendages, Anatomy, and Relationships of Trilobites
Raymond, Percy E. (Percy Edward)
Trilobites
The first thoracic limb is uniramous and similar to the maxillæ, but
the five following pairs are biramous swimming legs with coxopodite,
basipodite, exopodite, and endopodite. Both the exopodite and
endopodite are shorter than in the trilobites, but bear setæ and
spines.
The last pair of thoracic limbs are usually modified in the male into
copulatory organs. In some females they are enlarged to form plates
for the protection of the eggs, in others they are unmodified. In
still others they are much reduced or disappear. The abdomen is
without appendages.
The development in Copepoda is direct, by the addition posteriorly to
the larval form (nauplius) of segments, and the appendages remain
nearly unmodified in the adult.
Altogether, the primitive Copepoda seem much more closely allied to
the Trilobita than any other modern Crustacea, but unfortunately no
fossil representative of the subclass has been found. This is not so
surprising when one considers the habits and the habitat of most of
the existing species. Many are parasitic, many pelagic in both fresh
and marine waters, and many of those living on the bottom belong to
the deep sea or fresh water. Most free-living forms are minute, and
all have thin tests.
The eyes of copepods are of interest, in that they suggest the paired
ocelli of the Harpedidæ and Trinucleidæ. In the Copepoda there are, in
the simplest and typical form of these organs, three ocelli, each
supplied with its own nerve from the brain. Two of these are dorsal
and look upward, while the third is ventral. In some forms the dorsal
ocelli are doubled, so that five in all are present (cf. some species
of Harpes with three ocelli on each mound). In some, the cuticle over
the dorsal eyes is thickened so as to form a lens, as appears to be
the case in the trilobites. These peculiar eyes may be a direct
inheritance from the Hypoparia.
ARCHICOPEPODA.
Professor Schuchert has called my attention to the exceedingly curious
little crustacean which Handlirsch (1914) has described from the
Triassic of the Vosges. Handlirsch erected a new species, genus,
family, and order for this animal, which he considered most closely
allied to the copepods, hence the ordinal name. _Euthycarcinus
kessleri_, the species in question, was found in a clayey lens in the
Voltzia sandstone (Upper Bunter). Associated with the new crustacean
were specimens of _Estheria_ only, but in the Voltzia sandstone itself
land plants, fresh and brackish water animals, and occasionally,
marine animals are found. The clayey lens seems to have been of fresh
or brackish water origin.
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