The Appendages, Anatomy, and Relationships of TrilobitesRaymond, Percy E. (Percy Edward)
Science
The Appendages, Anatomy, and Relationships of Trilobites
Raymond, Percy E. (Percy Edward)
Trilobites
All of the specimens (three were found) are small, about 35 mm. long
without including the caudal rami, crushed flat, and not very well
preserved. The head is short, not so wide as the succeeding segments,
and apparently has large compound eyes at the posterior lateral
angles. The thorax consists of six segments which are broader than the
head or abdomen. The abdomen, which is not quite complete in any one
specimen, is interpreted by Handlirsch as having four segments in the
female and five in the male. Least satisfactory of all are traces of
what are interpreted by the describer as a pair of long stiff
unsegmented cerci or stylets on the last segment.
The ventral side of one head shield shows faint traces of several
appendages which must have presented great difficulty in their
interpretation. A pair of antennules appear to spring from near the
front of the lower surface, and the remainder of the organs are
grouped about the mouth, which is on the median line back of the
center. Handlirsch sees in these somewhat obscure appendages four
pairs of biramous limbs, antennæ, mandibles, maxillulæ, and maxillæ,
both branches of each consisting of short similar segments,
endopodites and exopodites being alike pediform.
Each segment of the thorax has a pair of appendages, and those on
the first two are clearly biramous. The endopodites are walking legs
made up of numerous short segments (twelve or thirteen according to
Handlirsch's drawing), while the exopodite is a long breathing and
rowing limb, evidently of great flexibility and curiously like the
antennules of the same animal. The individual segments are narrow at
the proximal end, expand greatly at the sides, and have a concave
distal profile. A limb reminds one of a stipe of _Diplograptus_.
Both branches are spiniferous.
No appendages are actually present on the abdomen, but each segment
has a pair of scars showing the points of attachment. From the small
size of these, it is inferred that the limbs were poorly developed.
This species is described in so much detail because, if it is a
primitive copepod, it has a very important bearing on the ancestry of
that group and is the only related form that has been found fossil.
The non-parasitic copepods have typically ten (eleven) free segments,
including the telson, and the four abdominal segments are much more
slender than the six in front of them. In this respect the agreement
is striking, and the presence of five pairs of appendages in the head
and six free segments in the thorax is a more primitive condition than
in modern forms where the first two thoracic segments are apparently
fused (Calman, 1909, p. 73).
The large compound eyes of this animal are of course not present in
the copepods, but as vestiges of eyes have been found in the young of
_Calanus_, it is possible that the ancestral forms had eyes.
Public-domain text, read in full here on John Shaqi.
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