The Appendages, Anatomy, and Relationships of TrilobitesRaymond, Percy E. (Percy Edward)
Science
The Appendages, Anatomy, and Relationships of Trilobites
Raymond, Percy E. (Percy Edward)
Trilobites
Burmeister (1843-1846), in his "Organization of the Trilobites,"
reviewed in _extenso_ the history of the search for appendages, and
concluded that they must have been so soft as to preclude the
possibility of their being preserved as fossils. "Their very absence
in fossils most distinctly proves their former real structure" (p.
10). In figures 7 and 8 on plate 6 he gave a restoration of the
ventral surface of an _Asaphus_, the first restoration of the ventral
anatomy to be attempted. Since he chose modern branchiopods as his
model, he did not go so far wrong as he might have done. Still, there
is little in the figure that would now be accepted as correct. The
following quotation will serve to give the opinion of this zoologist,
who from his knowledge of the Crustacea, was the most competent of the
men of his time to undertake a restoration of the appendages of the
trilobites:
... in giving a certain form to the feet in the restored figure, I
have done so rather intending to indicate what they might have
resembled, than with any idea of assuming their actual form. I
merely assert that these organs were soft, membranous, and fringed,
adapted for locomotion in water, placed on the abdominal portion of
the body, and extending sidewise beneath the lateral lobes of the
rings, as shown in the ideal transverse section. These feet were
also indented, and thus divided into several lobes at the open
lower side, and each separate lobe was furnished at the margin with
small bristles serving as fins. The last and external lobe was
probably longer, smaller, and more movable, and reached to the
termination of the projecting shell lobe, bearing a bladder-shaped
gill on the inner side (1846, p. 45).
McCoy (1846) observed in several trilobites a pair of pores situated
in the dorsal furrows near the anterior end of the glabella. He showed
that the pits occupy precisely the position of the antennæ of insects
and suggested that they indicated the former presence of antennæ in
these trilobites (chiefly _Anipyx_ and "_Trinucleus_"). The evidence
from _Cryptolithus_, set forth on a later page, indicates the
correctness of McCoy's view.
Richter (1848, p. 20, pl. 2, fig. 32) described and figured what he
took to be a phyllopod-like appendage found in a section through a
_Phacops_. Without the specimen it is impossible to say just what the
structure really was. The outline figure is so obviously modeled on an
appendage of _Apus_ that one is inclined to think it somewhat
diagrammatic. In calling attention to this neglected "find," Clarke
(1888, p. 254, fig.) interprets the appendage as similar to the spiral
branchiæ of _Calymene senaria_, and adds that he himself has seen
evidence of spiral branchiæ in the American Phacops rana.
Beyrich (1846) described a cast of the intestine of "_Trinucleus_,"
and Barrande (1852) further elaborated on this discovery.
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account