The Appendages, Anatomy, and Relationships of Trilobites — John Shaqi
The Appendages, Anatomy, and Relationships of TrilobitesRaymond, Percy E. (Percy Edward)
Science
The Appendages, Anatomy, and Relationships of Trilobites
Raymond, Percy E. (Percy Edward)
Trilobites
The first mention of _Neolenus_ with appendages preserved was in
Doctor Walcott's paper of 1911, in which two figures were given to
show the form of the exopodites in comparison with the branchiæ of the
eurypterid-like _Sidneyia_. In 1912, two more figures were presented,
showing the antennules, exopodites, and cerci. The specimens were
found in the Burgess shale (Middle Cambrian) near Field, in British
Columbia. This shale is exceedingly fine-grained, and has yielded a
very large fauna of beautifully preserved fossils, either unknown or
extraordinarily rare elsewhere. It was stated in this paper (1912 A)
that trilobites, with the exception of _Agnostus_ and _Microdiscus_,
were not abundant in the shale.
In discussing the origin of the tracks known as _Protichnites_,
Walcott presented four figures of _Neolenus_ with appendages, and
described the three claw-like spines at the tip of each endopodite.
Three new figures of the appendages were also contributed to the
second edition of the Eastman-Zittel "Text-book of Paleontology"
(1913, p. 701). Later (1916, pl. 9) there was published a photograph
of a wonderful slab, bearing on its surface numerous Middle Cambrian
Crustacea. Several of the specimens of _Neolenus_ showed appendages.
Finally, in 1918, appeared the "Appendages of Trilobites," in which
the limbs of _Neolenus_ were fully described and figured (p. 126),
and a restoration presented. Organs previously unknown in trilobites,
epipodites and exites, attached to the coxopodites, were found.
=Neolenus serratus= (Rominger).
(Text fig. 2-8.)
Illustrated: Walcott, Smithson. Misc. Coll., vol. 57, 1911, p. 20,
pl. 6, figs. 1, 2 (exopodites of thorax and cephalon);--Ibid., vol.
57, 1912, p. 191, pl. 24, figs. 1, la (antennules, caudal rami, and
endopodites of thorax);--Ibid., vol. 57, 1912, p. 277, pl. 45,
figs. 1-4 (antennules, endopodites of cephalon and thorax, caudal
rami);--Text-book of Paleontology, edited by C. R. Eastman, 2d ed.,
vol. 1. 1913, p. 701, fig. 1343 (exopodites), p. 716, fig. 1376
(abdominal appendages), fig. 1377 (appendages of thorax and
pygidium);--Ann. Rept. Smithson. Inst. for 1915, 1916, pl.
9;--Smithson. Misc. Coll., vol. 67, 1918, pp. 126-131 et al., pl.
14, fig. 1; pls. 15-20; pl. 21, fig. 6; pls. 22, 23; pl. 31
(restoration); pl. 34, fig. 3 (restored section); pl. 35, fig. 4;
pl. 36, fig. 3 (hypostoma).
The following description of the appendages of _Neolenus_ is
summarized from Walcott's paper of 1918, and from a study of the eight
specimens mentioned below.
_Cephalon._
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