The Appendages, Anatomy, and Relationships of TrilobitesRaymond, Percy E. (Percy Edward)
Science
The Appendages, Anatomy, and Relationships of Trilobites
Raymond, Percy E. (Percy Edward)
Trilobites
diagonally forward and outward. (pl. 21, fig. 6; pl. 22.)
_Pygidium._
The pygidium of _Neolenus serratus_ is large, and usually shows five
rings on the axial lobe and four pairs of ribs on the sides. There are
five pairs of biramous appendages belonging to this shield, and behind
these a pair of jointed cerci. That the number of abdominal appendages
should correspond to the number of divisions of the axial lobe rather
than to the number of ribs on the pleural lobes is of interest, and in
accord with other trilobites, as first shown by Beecher.
The endopodites of the pygidium have the same form as those of the
thorax, are long, and very much less modified than those of any other
trilobite whose appendages are known. On some specimens, they extend
out far beyond the dorsal test, so that nearly all the segments are
visible (pl. 17, fig. 3; pl. 18; pl. 19; pl. 20, fig. 1), but in these
cases are probably displaced. The segments are short and wide, the
whole endopodite tapering gradually outward. The dactylopodite bears
terminal spines, and the individual segments also have outward-directed
spines.
The cerci appear to have been long, slender, very spinose organs much
like the antennules, but stiff rather than flexible. They are a little
longer than the pygidium (pl. 17, figs. 1, 2), and seem to be attached
to a plate on the under surface of the posterior end and in front of
the very narrow doublure. The precise form of this attachment can not
be determined from the published figures. They bear numerous fine
spines (pl. 17, fig. 3).
_Epipodites and Exiles._
Doctor Walcott has found on several specimens of _Neolenus_ remains of
organs which he interprets as epipodites and exites attached to the
coxopodites. A study of the specimens has, however, convinced me that
both the large and small epipodites are really exopodites, and that
the exites are badly preserved and displaced coxopodites. Detailed
explanation of this interpretation is given below in the description
of the several specimens involved.
_Description of Individual Specimens._
Doctor Walcott was kind enough to send me eight of the more important
specimens of _Neolenus_ figured by him, and since my interpretation
of them does not agree in all respects with his, I have thought it
fairer to the reader to present here rather full notes explaining
the position I have taken. I understand that since I communicated my
interpretation of the epipodites and exites to him, Doctor Walcott has
submitted the specimens to several palæontologists, who consider that
epipodites are really present. Since I am not able to convince myself
that their conclusion is based upon sound evidence, I give here my own
interpretation. There is of course, no a priori reason why trilobites
should not have had epipodites.
Specimen No. 58589.
Public-domain text, read in full here on John Shaqi.
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