The Appendages, Anatomy, and Relationships of Trilobites — John Shaqi
The Appendages, Anatomy, and Relationships of TrilobitesRaymond, Percy E. (Percy Edward)
Science
The Appendages, Anatomy, and Relationships of Trilobites
Raymond, Percy E. (Percy Edward)
Trilobites
In the same year Woodward (1884, p. 162, fig. 1-3) reproduced all of
Mickleborough's figures, and suggested that the last seven pairs of
appendages on the pygidium of _Calymene_ and _Isotelus_ were probably
"lamelliform branchiferous appendages, as in _Limulus_ and in living
Isopoda."
Professor Beecher published, in 1902, an outline taken from
Mickleborough's figure of this specimen, to call attention to certain
discontinuous ridges along the axial cavity of the anterior part of
the pygidium and posterior end of the thorax. These ridges are well
shown in Mickleborough's figure, though not in that of Walcott, and
their presence on the specimen was confirmed by a study by Schuchert,
who contributed a diagrammatic cross-section to Beecher's paper (1902,
p. 169, pl. 5, figs. 5, 6). Beecher summarized in a paragraph his
interpretation of this specimen:
The club-shaped bodies lying within the axis are the gnathobases
attached at the sides of the axis; the curved members extending
outward from the gnathobases are the endopodites; the longitudinal
ridges in the ventral membrane between the inner ends of the
gnathobases are the buttresses and apodemes of the mesosternites;
the slender oblique rod-like bodies shown in the right pleural
region in Walcott's figure are portions of the fringes of the
exopodites.
In 1910, Mr. W. C. King of Ottawa, Ontario, found at Britannia, a few
miles west of Ottawa, the impression in sandstone of the under surface
of a large specimen of _Isotelus arenicola_, described on a later page
(p. 39).
Finally (1918, p. 133, pl. 24, figs. 3, 3a; pl. 25), Walcott has
redescribed the specimen from Ohio, presenting a new and partially
restored figure. He refers also to the specimen from Ottawa under the
name _Isotelus covingtonensis?_ Foerste (not Ulrich). He advances the
view, which I am unable to share, that the cylindrical appearance of
the segments of the appendages of _Isotelus_ is due to post-mortem
changes.
=Isotelus latus= Raymond.
(pl. 10, fig. 1.)
Illustrated: _Asaphus platycephalus_ Billings, Quart. Jour. Geol.
Soc., London, vol. 26, 1870, pl. 31, figs. 1-3; pl. 32, figs. 1,
2.--Woodward, Geol. Mag., vol. 8, 1871, pl. 8, figs. 1,
1a.--Gerstäcker, in Bronn's "Klassen u. Ordnungen d. Thier-Reichs,"
1879, pl. 49, fig. 1.--von Koenen, N. Jahrb. f. Min., etc., vol. 1,
1880, pl. 8, fig. 8.--Milne-Edwards, Ann. Sci. Nat., Zoologie, ser.
6, vol. 12, 1881, pl. 12, fig. 45.
_Isotelus latus_ Raymond, Bull. Victoria Mem. Mus., Geol. Survey
Canada, No. 1, 1913, p. 45 (species named).
_Isotelus covingtonensis?_ Walcott (not Foerste), Smithson. Misc.
Coll., vol. 67, 1918, p. 134.
Knowledge of the appendages of this species is derived from the
specimen which Billings described in 1870. It was found in the
Trenton, probably the Middle Trenton, near Ottawa, Ontario, and is
preserved in the Victoria Memorial Museum at Ottawa.
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