The Appendages, Anatomy, and Relationships of TrilobitesRaymond, Percy E. (Percy Edward)
Science
The Appendages, Anatomy, and Relationships of Trilobites
Raymond, Percy E. (Percy Edward)
Trilobites
THE APPENDAGES OF PTYCHOPARIA.
=Ptychoparia striata= (Emmrich).
Illustrated: Jaekel, Zeits. d. d. geol. Gesell., 1901, vol. 53,
part 1, pls. 4, 5.
Jaekel has described a specimen of this species obtained from the
Middle Cambrian near Tejrovic, Bohemia, which on development showed
beneath the test of the axial lobe, certain structures which he
believed represented the casts of proximal segments of appendages.
On the basis of this specimen he produced a new restoration of the
ventral surface of the trilobite, in which he showed three short wide
segments in the place occupied by the coxopodite of an appendage of
_Triarthrus_. He also made the mouth parts considerably different from
those of the latter genus. Beecher (1902) showed that the structures
which Jaekel took for segments of appendages were really the fillings
between stiffening plates of chitin on the ventral membrane, and
demonstrated the fact that similar structures existed in _Triarthrus_.
It cannot be said, therefore, that any appendages are really known in
_Ptychoparia striata_, but some knowledge of the internal anatomy of
the species is supplied by the specimen.
=Ptychoparia cordilleræ= (Rominger).
Illustrated: Walcott, Smithson. Misc. Coll., vol. 57, 1912, p. 192,
pl. 24, fig. 2;--Ibid., vol. 67, 1918, pl. 21, figs. 3-5 (corrected
figure).
Walcott has figured a single individual of this species showing
appendages, the accompanying description being as follows (1918, p.
144):
Ventral appendages. Only one specimen has been found showing the
thoracic limbs. This indicates very clearly the general character
of the exopodite and that it is situated above the endopodite,
although there are only imperfect traces of the latter....
The exopodites are unlike those of any trilobite now known. They
are long, rather broad lobes extending from the line of the union
of the mesosternites and the pleurosternites. At the proximal end
they appear to be as wide as the axial lobe of each segment, and to
increase in width and slightly overlap each other nearly out to the
distal extremity.... They are finely crenulated along both the
anterior and dorsal margins, which indicates the presence of fine
setæ.
The specimen is quite imperfectly preserved, but seems to indicate
that the exopodite of Ptychoparia had a long, rather narrow
unsegmented shaft.
_Measurements_ (from Walcott's figure): The specimen is a small one,
about 9.5 mm. long, an individual exopodite is about 2 mm. long and
the shaft 0.33 mm. wide.
_Horizon and locality:_ Middle Cambrian, Burgess shale, between Mount
Field and Wapta Peak, above Field, British Columbia.
=Ptychoparia permulta= Walcott.
Illustrated: Walcott, Smithson. Misc. Coll., vol. 67, 1918, p. 145,
pl. 21, figs. 1, 2.
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