The Appendages, Anatomy, and Relationships of TrilobitesRaymond, Percy E. (Percy Edward)
Science
The Appendages, Anatomy, and Relationships of Trilobites
Raymond, Percy E. (Percy Edward)
Trilobites
Now consider the "epipodites." They are well shown in _Calymene_ in
the specimens illustrated on plate 27, figure 11 (1918), and plate 3,
figure 3 (1881), and less clearly in one or two others. Slices 22 (pl.
27, fig. 12, 1918) and 80 (our fig. 12) show what is called the same
organ in Ceraurus. It will be noted that all of these slices are cut
in the same way, that is, more or less parallel to the under surface
of the head, or, at any rate, on a plane parallel to a plane which
would be tangent to the axial portion of the coiled shell. The
sections which show the spirals best are those which are cut by a
plane perpendicular to the long axis of the body. If one were to
attempt to cut an enrolled _Triarthrus_ in such a way as to get a
section showing the length of the setæ, one would not cut a section
perpendicular to the axis of the animal, nor, in fact, would he cut
one parallel to the ventral plane, but it is obvious that in this
latter type of section he would stand a better chance of finding a
part of the plane of the exopodite coincident with the plane of his
section than in the former. And that seems to be what has happened in
these sections of _Calymene_ and _Ceraurus_. If the exopodites were
preserved, transverse sections were bound to cut across many sets of
fringes, and the resultant slice would show transverse sections of the
setæ as a series of overlapping spots. A few fortunately located
sections in a more nearly horizontal plane might cut the setæ and
occasionally the shaft of one or more exopodites in the longitudinal
plane, and the resulting effect would produce the so-called
"epipodites." A careful study has shown that no one of these
epipodites is complete, and they do not have the palmate form shown in
Walcott's figures.
And the last and most important argument against the spiral appendages
is that certain slices, of both _Calymene_ and _Ceraurus_, show
definitely exopodites of exactly the type found in other trilobites.
These are discussed later in the detailed description of the various
slices.
If these series of spots are interpreted on the basis of the known
structure of _Triarthrus_, they are of course a series of sections
through the setæ of the exopodites. It will be shown in Part IV
that these setæ are not circular in section, but flattened, in
_Cryptolithus_ even blade-like, and that they overlap one another. A
section across them would give the same general appearance as, for
instance, that shown in figures 4, 6, 9, and 10 of Walcott's plate 3
(1881).
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