Wynne-Edwards (1935:241) defines the offshore zone as extending to the
continental edge; however, in Micronesia where small islands rise
abruptly out of the ocean's depths, there is no useful way to separate
the offshore zone from the pelagic zone. Since certain species go
farther from the land than others, the two zones may be combined as a
single zone extending beyond the sight of land. Birds which frequent
this area beyond the inshore zone but may not range extensively at sea
include _Fregata_, _Sula_, _Sterna fuscata_, _S. hirundo_, _S.
anaetheta_, and others. The Herring Gull (_Larus argentatus_), which has
been taken in the northern Marianas, may be classed with this group
although it probably ranges widely in the open sea. Birds which spend
considerable time at sea and may seldom approach land include _Diomedea
nigripes_, the petrels (_Puffinus_ and _Pterodroma_), and possibly the
tropic birds (_Phaëthon_).
In numbers of individuals the birds inhabiting the inshore zones are
relatively more numerous than those preferring the offshore and pelagic
zones, although 12 of the 18 resident kinds of oceanic birds apparently
prefer the offshore zone, while only 6 kinds appear to be restricted
primarily to the inshore areas.
FAUNAL COMPONENTS
The oceanic birds were probably among the earliest birds to reach the
islands of Micronesia. The presence of phosphate deposits on islands
(Fais, Angaur), denoting deposition of guano by oceanic birds (possibly
boobies, noddies, sooty terns), indicates long time residence by these
birds. A person is prone to think that these deposits must have been
made by larger concentrations of oceanic birds than are found in these
islands today. Whether there were actually more individuals present
during the period of deposition of phosphate in the lagoons of these
islands is not known, although the elevation of the lagoons (forming the
raised islands of Fais and Angaur) with the resulting freshening of the
water probably was a great attraction to oceanic birds, especially to
those which prefer to drink fresh water. According to Leonard P. Schultz
(_in litt._), the abundance of fish in the areas about these Pacific
islands has been approximately the same since Pleistocene times, so that
there was apparently no greater concentration of fish near these islands
to attract large populations of fish-eating sea birds. Probably the time
element is of sufficient magnitude to account for such deposition by
birds with a population similar to that found there today.
The oceanic avifauna of Micronesia contains birds which are apparently
from ancestral homes in the Palearctic Region, in the North and Central
Pacific, in Polynesia, in Melanesia and Malaysia, and from homes the
positions of which are uncertain because of the widespread
circumtropical occurrence of the birds. There are no sea birds that are
endemic in Micronesia.
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