Mayr (1941b:203) cites _Psamathia_ as an example of "restricted
endemism" and points out that the nearest relative occurs in the
Philippines. Delacour (1942:514), in a discussion of the bush-warblers
of the genera _Cettia_, _Bradypterus_ and related forms, says,
"_Psamathia annae_, from Palau Islands, is related to _Cettia_,
differing mainly in its much longer bill and legs."
_Psamathia_ is a specialized bush-warbler and has followed a pattern of
evolution which characterizes some of the other island birds in that the
bill and legs are long and the wing is rather short and rounded.
_Psamathia_ resembles many of the bush-warblers, as well as the
reed-warblers {_Acrocephalus_); in general, body coloring being paler
below and darker above. It differs from _Acrocephalus_ by having a
longer tenth primary, smaller second and third primaries, only ten tail
feathers, a more rounded wing, differently shaped nostrils, and by much
softer plumage (the latter character is found also in _Collurcincla
tenebrosus_ and _Cleptornis marchei_ of Micronesia). Rather than being
related to the reed-warblers, as was supposed by Hartlaub and Finsch,
_Psamathia_ seems closest to _Cettia_, especially to _Cettia_
(_Horeites_) _diphone seebohmi_ of the Philippine Islands. _Psamathia_
has a longer bill than this bird, but the general appearance and
structure of the feet, tail, wing, body and bill are the same.
=Acrocephalus luscinia luscinia= (Quoy and Gaimard)
Nightingale Reed-warbler
_Thryothorus luscinius_ Quoy and Gaimard, Voy. "l'Astrolabe,"
Zool., 1, 1830, p. 202, pl. 5, fig. 2. (Type locality, Marian Is. =
Guam.)
_Sylvia syrinx_ Kittlitz (part), Obser. Zool., in Lutké, Voy. "Le
Seniavine," 3, 1836, p. 306 (Guahan); _idem_ (part), Denkw. Reise
russ. Amer. Micron. und Kamchat., 2, 1858, p. 141 (Guaham).
_Tatare luscinia_ Gray, Genera Birds, 3, 1849, App. 8 (Marian Is.
= Guam); Hartlaub, Journ. f. Ornith., 1854, p. 167 (Mariannen =
Guam); Gray, Cat. Birds Trop. Is. Pacific Ocean, 1859, p. 14
(Ladrone or Marian Is. = Guam); Finsch, Journ. Mus. Godeffroy, 12,
1876, p. 31 (Guaham); Wiglesworth, Abhandl. und Ber. Zool. Mus.
Dresden, no. 6, 1890-1891 (1891), p. 41 (Guam); Büttikofer, Notes
Leyden Mus., 14, 1892, p. 16 (Guam); Oustalet, Nouv. Arch. Mus.
Hist. Nat. Paris, (3), 7, 1895, p. 209 (Guam).
_Tatare luscinius_ Bonaparte, Consp. Avium, 1, 1850, p. 224 (Guam);
_idem_, Comptes Rendus Acad. Sci. Paris, 41, 1855, p. 1111
(Mariannes = Guam); Gray, Hand-list Birds, 1, 1869, p. 194 (Ladrone
= Guam).
_Hybristes_ [_luscinia_] Reichenbach, Syst. Avium, 1850, pl. 57,
fig. 7 (no locality = Guam).
_Acrocephalus orientalis_ Pelzeln, Reise, "Novara," Vögel, 1865,
p. 64 (Guaham).
_Tatares luscinius_ Giebel, Thes. Ornith., 3, 1877, p. 599
(Marianae).
_Acrocephalus mariannae_ Tristram, Ibis, 1883, p. 45 (Type
locality, Guam).
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