The well ossified lateral processes and enlarged base of Didier's
(1954:240) specimen suggest that it is of a more mature individual than
the one described above. These specimens of _M. nivalis_, as well as the
specimens of _M. longicaudus_, exhibit dorso-ventral flattening of the
mid-part of the base of the stalk.
The baculum of a specimen from Switzerland is weakly developed, of small
size (shaft 2.0 mm. in length), slender, thin, spatulate, and terminally
inflated. Digital processes were not observed, perhaps owing to excessive
maceration in preparation. The general appearance of the baculum is that
of an immature individual, although the animal was not small (165 mm.
total length in preservative).
_Specimens examined_: Two _Microtus nivalis nivalis_; Zermatt, Valais,
Switzerland, 67105; Wetterstein, Germany, 65127.
Microtus (Chionomys) longicaudus (Merriam)
Fig. 48
Baculum: Base of stalk well developed, greatest length (3 mm.) 1-3/4
times greatest breadth, 3-2/3 times greatest depth; three ossified
processes; base of median process rounded; median process slightly curved
in dorsoventral plane, in length almost 1/3 greatest length of stalk;
ossifications in lateral processes variable in size, frequently widely
separated from shaft by cartilage, rarely as large as median
ossification; basal tuberosities usually well-developed, medially
confluent; profile of base in dorsal view trilobate or irregularly convex
throughout; constriction 1/2 greatest depth; shaft relatively straight or
slightly bowed ventrally or dorsally, shaft at mid-point of stalk wider
than high; tip of shaft laterally inflated; widest point of stalk
approximately 1/4 length of stalk from proximal end; lateral profile in
dorsal view tapers gradually onto shaft anteriorly from point of greatest
width of stalk; shaft variable, from slender terminally and nearly
parallel sided (Fig. 48), to broad distally and tapered.
In many of the features that distinguish _M. longicaudus_ (and the
closely related insular species _M. coronarius_) from other North
American _Microtus_, _longicaudus_ resembles the Old World species of the
subgenus _Chionomys_ (that is to say, _M. nivalis_, _M. gud_, and _M.
roberti_). These features are medium size, long tail, grayish color,
montane habitat, relatively short molar tooth-row, moderate sized and
unconstricted incisive foramen, relatively decurved upper incisors,
elongate nasals, relatively broad interorbital region without
well-developed median ridge, and similar chromosomes (Matthey, 1955:178).
For these reasons I am here referring _Microtus longicaudus_ to the
subgenus _Chionomys_; previously it has not been referred to that
subgenus.
_Specimens examined_: Six, of three subspecies; _Microtus longicaudus
littoralis_, Sullivan Island, Alaska, 42972, 42969; _M. l. mordax_, 3/4
mi. N, 2 mi. W Allenspark, 8400 ft., Boulder Co., Colorado, 50335, 76829;
_M. l. sierrae_, Crane Flat, Mariposa Co., California, 50252, 50253.
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