17´. Length of stalk usually less than 3.6 mm., or if more than
3.6 mm. (up to 4.0 mm.) then length 1-1/2 times or more its
greatest breadth 18
18. Median process attenuate distally in dorsal view, and
relatively long (more than twice its own breadth), 1/5 to 3/5 the
length of stalk; breadth of stalk usually 2/3 or more length of
stalk.... _Microtus miurus_, p. 200
18´. Median process relatively blunt distally in dorsal view,
relatively short (usually less than 1/4 length of stalk), breadth
of stalk usually less than 2/3 length of stalk....
_Pitymys_, p. 208, _Pedomys_, p. 207, or _Microtus mexicanus_, p. 205
19. Distal processes small and firmly ankylosed to distal end of
shaft.... _Phenacomys longicaudus_, p. 197
19´. Distal processes if present not firmly ankylosed to distal
end of shaft 20
20. Dorsal concavity of base as viewed from proximal end usually
deeper than ventral concavity.... _Microtus mexicanus_, p. 205
20´. Dorsal and ventral concavities of base equal in depth or
ventral one the deeper 21
21. Total length of baculum more than 3.6 mm.... _Microtus
californicus_, p. 205
21´. Total length of baculum less than 3.6 mm.... _Synaptomys
cooperi_, p. 194
DISCUSSION
Owing to shortness of lower incisors and present geographic distribution
of the species, Hinton (1926:35) considered the Tribe Lemmi (lemmings) to
be more primitive than the Tribe Microti (voles). The surviving lemmings
are specialized in many features and therefore are considered as advanced
end-products of an evolutionary radiation of a primitive microtine stock,
of which all earlier stages are extinct.
Hinton regarded _Dicrostonyx_ as the most primitive of the genera of
lemmings on account of its more complex molar teeth (complexity was
considered to be primitive), and on account of the presence of three
primitive longitudinal rows of tubercles in unworn molars. The other
three genera were arranged in order of increasing specialization as
follows: _Synaptomys_, _Myopus_, _Lemmus_.
Public-domain text, read in full here on John Shaqi.
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