The Biological Problem of To-day: Preformation Or Epigenesis?: The Basis of a Theory of Organic DevelopmentHertwig, Oscar
Science
The Biological Problem of To-day: Preformation Or Epigenesis?: The Basis of a Theory of Organic Development
Hertwig, Oscar
Developmental biology; Embryology; Genetics
Weismann[2] leads the van for preformation; for the last ten years he has
occupied himself with the theoretical discussion of the questions set forth
above; and now, in a recent treatise, _The Germplasm_, he has combined his
views, already many times modified, in a coherent theory. Now he explains
candidly that he has been driven to the view that epigenetic development
does not exist. 'In the first chapter of my book,' he remarks, 'will be
found an actual proof of the reality of evolution, a proof so simple and
obvious that I can scarcely understand to-day how it could have escaped my
notice so long' (_Germplasm_, p. 14). Elsewhere he writes: 'I believe that
I have established that ontogeny can be explained only by evolution, and
not by epigenesis.'
A mental process, which consciously or unconsciously plays a great part
with evolutionists, and helps to determine their conclusions, is
characteristic of the direction of their inquiries. They set out from the
fact that the characters of the parents, often to the smallest detail, are
transmitted to children by means of the germ or rudiment; they conclude
that the active causes of all the complexity that arises must be contained
in the apparently homogeneous germ, embryological differentiation being a
spontaneous process. It follows that the apparent homogeneity is, in
reality, latent complexity which becomes patent during the progress of
ontogeny. Latent complexity implies a material substratum, consisting of
actual particles for which many different names have been found. As our
senses can give us no experimental knowledge of these particles, which are
so small as to be invisible, modern evolutionists attempt to picture them,
in imagination, by reflecting all the visible characters of the perfected
organism upon the undivided egg-cell, so peopling that globule of yolk with
a system of minute particles corresponding in quality and in spacial
arrangement with the larger parts of the adult.
Weismann has practised this art in the true spirit of a virtuoso, and has
elaborated it into a novel mode of biological investigation. Take an
example;--'It would be impossible,' he says in _The Germplasm_ (p. 138),
'for any small portion of the human skin to undergo a hereditary and
independent change from the germ onwards, unless a small vital element
corresponding to this particular part of the skin existed in the germ
substance, a variation in this element causing a corresponding variation in
the part concerned. Were this not the case, birth-marks would not exist.'
Public-domain text, read in full here on John Shaqi.
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