The Biological Problem of To-day: Preformation Or Epigenesis?: The Basis of a Theory of Organic DevelopmentHertwig, Oscar
Science
The Biological Problem of To-day: Preformation Or Epigenesis?: The Basis of a Theory of Organic Development
Hertwig, Oscar
Developmental biology; Embryology; Genetics
The biophores are his smallest material units, and to them are due the
fundamental qualities of life--assimilation, metabolism, and reproduction
by division. Thus, they correspond to Herbert Spencer's physiological
units, Darwin's gemmules, De Vries' pangenes, and Hertwig's idioblasts.
They are the bearers of the various characters of cells, and there are
present in the germplasm a very large multitude of different kinds of them,
corresponding to the number of cells with different characters.
The determinants are units of the rank next higher; they have qualities of
their own, but are composed of groups of several kinds of biophores. They,
too, have the power of division which is associated with, and comes about
by, multiplication of the coherent company of biophores which lies within
them.
The histological character of every cell in a multicellular organism is
determined by a single determinant (cell-determinants). Weismann has framed
his conception of determinants so as to avoid the supposition that every
single cell is represented in the germplasm by its own biophores. There are
small parts in the body in which the cells are all alike, and for these
parts a single determinant suffices, afterwards multiplying by division. On
the other hand, each cell or cell-group in the body, that is independently
variable, must have its special determinant in the germplasm. And so the
germplasm of a species must possess as many determinants, or guiding
particles, as there are in the organism cells or cell-groups that are
independently variable in the germ or in later stages (hereditary pieces or
determinates).
As every cell or group of cells which corresponds to determinants has a
definite position in the body, Weismann infers that the determinants are
definitely placed in the germplasm, and form an ordered, complicated
community. He has given the name id to these communities, which are higher
units with definite constitution and with complicated architecture. These
_ids_ are bodies containing all the determinants necessary to build up the
individual of a species, and correspond to what Weismann previously called
ancestral plasms. Every id must be able to grow and multiply, for it is by
their multiplication that the germplasm for new individuals is formed.
A single _id_ would suffice for the conduct of a single life-history;
Weismann, however, in the pursuit of a chain of thought connected with the
relation of sexual reproduction to heredity, and which I shall not discuss
here, regards the germplasm as being still more complicated, and consisting
of many, sometimes more than a hundred, ancestral plasms or _ids_, which
have been derived from near or distant ancestors, the peculiarities of
whose structures they retain, and may at some time actually produce
(explanation of atavism).
Public-domain text, read in full here on John Shaqi.
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