The approved classification of flies is to some extent dependent upon
the formation of the antennæ, but the unique feature of the systematic
differentiation is based upon a very intricate method of scrutinising,
identifying, and numbering the vein-like strengthening ribs called
veins, nervures, or nerve-lines, which, starting from the shoulder, mark
with characteristic patterns the transparent tissue of the wing. We are
rather compelled to follow something like this plan (simplified) for the
purpose of clearly distinguishing the "lesser house-fly" from the common
"house-fly."
In the accompanying illustrations rather similar patterns of wings are
shown; these are typical of the _Muscidæ_ and _Anthomyidæ_, which, taken
together, comprise amongst others all the cattle and human pestering
"sweat-flies"; only a few really blood-sucking flies are included
amongst the _Muscidæ_.
In critically comparing these four patterns, the chief feature to be
observed is the small rib-like nervure called the "discal" "cross-vein,"
which is situate in the very middle of the wing, and which connects the
lowest of a group of longitudinal nerve-lines or veins in the front (or
upper) half of the wing to the uppermost of the other group of
longitudinal nerve-lines in the hind (or lower) half of the wing. Three
"main" longitudinal lines, technically termed "veins," are theoretically
recognised as constituting the upper group and four "main" longitudinal
lines the lower group; but these "veins" (numbered 1, 2, 3, 4, 5, 6 and
7) will be found to be varied in different families and genera, each
often with characteristic diverging branches, whilst some veins may be
rather inconspicuous or quite absent. We will here devote our attention
only to two such "veins," those respectively termed "vein 3" and "vein
4" which are connected in the very middle of the wing, as above
mentioned, by the small but always distinct "discal" "cross-vein." The
illustrated patterns herewith show wings divided into about twelve
compartments or cells, to all of which learned entomological writers
give troublesome technical names, not nearly so intelligible as the
nomenclature symbols of the late Rev. W. J. Wingate, explained in the
Appendix herewith. For present purposes a simple observation of the
(externomedial) vein "V, 4," where it is the lower boundary of the
(subapical) cell "O, 4^2," will suffice.
[Illustration: Fig. 3. Wing patterns contrasted.]
The pattern of the first figure illustrates the wing of the common
blue-bottle; here "vein 4" does not run at all straight in the last part
of its course, but curiously bends very suddenly upwards at an angle and
meets the margin very near to "vein 3." In the wing of a large
blue-bottle it will be easy to recognise this plan.
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