in the air passages, as in the simple reflex act of coughing or in vocalisation, the glottis must be closed by approximation of the vocal cords, and this is effected by a group of muscles termed the adductors, which pull on the pyramid cartilages in such a way that the vocal processes are drawn towards one another in the manner shown in fig. 7. Besides the abductor and adductor groups of muscles, there is a muscle which acts in conjunction with the adductor group, and by its attachments to the shield cartilage above and the ring cartilage below makes tense the vocal cords (_vide_ fig. 5); it is of interest to note that this muscle has a separate nerve supply to that of the abductor and adductor muscles. [Illustration: FIG. 5 Diagram after Testut (modified), showing the larynx from the front.] [Illustration: FIG. 6 Diagram after Testut (modified), showing the posterior view of the larynx with the muscles.] On the top of the pyramid cartilages, in the folds of mucous membrane which cover the whole inside of the larynx are two little pieces of yellow elastic cartilage; and in the folds of mucous membrane uniting these cartilages with the leaf-like lid cartilage (epiglottis) is a thin sheet of muscle fibres which acts in conjunction with the fibres between the two pyramid cartilages (_vide_ fig. 8). I must also direct especial attention to a muscle belonging to the adductor group, which has another important function especially related to vocalisation: it is sometimes called the vocal muscle; it runs from the pyramid cartilage to the shield cartilage; it apparently consists of two portions, an external, which acts with the lateral ring-shield muscle and helps to approximate the vocal cords; and another portion situated within the vocal cord itself, which by contracting shortens the vocal cord and probably allows only the free edge to vibrate; moreover, when not contracting, by virtue of the perfect elasticity of muscle the whole thickness of the cord, including this vocal muscle, can be stretched and thrown into vibration (_vide_ fig. 8). In the production of chest notes the whole vocal cord is vibrating, the difference in the pitch depending upon the tension produced by the contraction of the tensor (ring-shield) muscle. When, however, the change from the lower to the upper register occurs, as the photographs taken by Dr. French and reproduced in a lecture at the Royal Institution by Sir Felix Semon show, the vocal cords become shorter, thicker, and rounder; and this can be explained by supposing that the inner portion of the vocal muscle contracts at the break from the lower to the upper register (_vide_ fig. 11); and that as a result only the free edges of the cords vibrate, causing a change in the quality of the tone. As the scale is ascended the photographs show that the cords become longer and tenser, which we may presume is due to the continued action of the tensor muscle. Another explanation is possible, viz. that in