In the following pages on the Relation of the Brain to the mechanism of the Voice in Speech and Song, I intend, as far as possible, to explain the mechanism of the instrument, and what I know regarding the cerebral mechanism by which the instrument is played upon in the production of the singing voice and articulate speech. Before, however, passing to consider in detail the instrument, I will briefly direct your attention to some facts and theories regarding the origin of speech. THEORIES ON THE ORIGIN OF SPEECH The evolutionary theory is thus propounded by Romanes in his "Mental Evolution in Man," pp. 377-399: "Starting from the highly intelligent and social species of anthropoid ape as pictured by Darwin, we can imagine that this animal was accustomed to use its voice freely for the expression of the emotions, uttering danger signals, and singing. Possibly it may also have been sufficiently intelligent to use a few imitative sounds; and certainly sooner or later the receptual life of this social animal must have advanced far enough to have become comparable with that of an infant of about two years of age. That is to say, this animal, although not yet having begun to use articulate signs, must have advanced far enough in the conventional use of natural signs (a sign with a natural origin in tone and gesture, whether spontaneously or intentionally imitative) to have admitted of a totally free exchange of receptual ideas, such as would be concerned in animal wants and even, perhaps, in the simplest forms of co-operative action. Next I think it probable that the advance of receptual intelligence which would have been occasioned by this advance in sign-making would in turn have led to a development of the latter--the two thus acting and reacting on each other until the language of tone and gesture became gradually raised to the level of imperfect pantomime, as in children before they begin to use words. At this stage, however, or even before it, I think very probably vowel sounds must have been employed in tone language, if not also a few consonants. Eventually the action and reaction of receptual intelligence and conventional sign-making must have ended in so far developing the former as to have admitted of the breaking up (or articulation) of vocal sounds, as the only direction in which any improvement in vocal sign-making was possible." Romanes continues his sketch by referring to the probability that this important stage in the development of speech was greatly assisted by the already existing habit of articulating musical notes, supposing our progenitors to have resembled the gibbons or the chimpanzees in this respect. Darwin in his great work on the "Expression of the Emotions" points to the fact that the gibbon, the most erect and active of the anthropoid apes, is able to sing an octave in half-tones, and it is interesting to note that Dubois considers his Pithecanthropus Erectus is on the same stem as the gibbon. But it has