The Cambridge natural history, Vol. 02 (of 10)Beddard, Frank E. (Frank Evers)
Science
The Cambridge natural history, Vol. 02 (of 10)
Beddard, Frank E. (Frank Evers)
Animals
PROSTHIOSTOMATIDAE. {
Tentacles absent. Body {
elongated. Pharynx {_Prosthiostomum_ (Fig. 8, F).
long, cylindrical. {
Penis with accessory {
muscular vesicles. {
{20}APPEARANCE AND SIZE OF POLYCLAD TURBELLARIA.—Polyclads are almost
unique amongst animals in possessing a broad and thin, delicate body that
glides like a living pellicle over stones and weeds, moulding itself on to
any inequalities of the surface over which it is travelling, yet so fragile
that a touch of the finger will rend its tissues and often cause its speedy
dissolution. The dorsal surface in a few forms is raised into fine
processes (_Planocera villosa_), or into hollow papillae (_Thysanozoon
brocchii_), and in very rare cases may be armed with spines (_Acanthozoon
armatum_,[26] _Enantia spinifera_); in others, again, nettle-cells
(nematocysts) are found (_Stylochoplana tarda_, _Anonymus virilis_). Some
Polyclads, especially the pelagic forms, are almost transparent; in others,
the colour may be an intense orange or velvety black, and is then due to
peculiar deposits in the epidermal cells. Between these two extremes the
colour is dependent upon the blending of two sources, the pigment of the
body itself and the tint of the food. Thus a starved _Leptoplana_ is almost
or quite white, a specimen fed on vascular tissue reddish. Many forms are
coloured in such a way as to make their detection exceedingly difficult,
but this is probably not merely due, as Dalyell supposed, to the substratum
furnishing them with food and thus colouring them sympathetically, but is
probably a result of natural selection.
The largest Polyclad, the bulkiest Turbellarian, is _Leptoplana gigas_ (6
inches long and 4 in breadth), taken by Schmarda, free-swimming, off the
coast of Ceylon. The largest European form is _Pseudoceros maximus_, 3½
inches in length and stoutly built. A British species, _Prostheceraeus
vittatus_, attains a length of from 2 to 3 inches. These large forms,
especially the Pseudoceridae (pre-eminently the family of big Polyclads),
are brightly coloured, and usually possess good swimming powers, since,
being broad and flat, they are certainly not well adapted for creeping
rapidly, and this is well shown by the way these Polyclads take to swimming
when in pursuit of prey at night. The size of any individual is determined,
amongst other factors, by the period at which maturity sets in, after which
probably no increase takes place. Polyclads apparently live about twelve
months, and mature specimens of the same species vary from ½ inch to 2½
inches in length (_Thysanozoon brocchii_), {21}showing that growth is,
under favourable conditions, very rapid.
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