The Cambridge natural history, Vol. 02 (of 10) — John Shaqi
The Cambridge natural history, Vol. 02 (of 10)Beddard, Frank E. (Frank Evers)
Science
The Cambridge natural history, Vol. 02 (of 10)
Beddard, Frank E. (Frank Evers)
Animals
The Trematodes[2] may be divided into those living on the outer surface of
various aquatic animals, usually fish (Ectoparasites); and those which
penetrate more or less deeply into the alimentary canal or the associated
organs of the host (Endoparasites). They are oval, flattened
Platyhelminthes ranging from a microscopic size to a length of three feet
(_Nematobothrium_, Fig. 22), and are provided with organs of adhesion by
which they cling to the outer surface, or to the interior, of the animals
they inhabit. Trematodes occur parasitically in all groups of Vertebrates,
but, with the exception of the liver-flukes of the sheep (_Distomum
hepaticum_ and _D. magnum_), and of _Bilharzia haematobia_ found in man (in
the blood-vessels of the urinary bladder) over the greater part {5}of
Africa, their attacks are not usually of a serious nature. Ectoparasitic
Trematodes are _Monogenetic_; that is, their larvae grow up directly into
mature forms. The Endoparasitic species, however, are usually _Digenetic_.
Their larvae enter an Invertebrate and produce a new generation of
different larvae, and these another. The last are immature flukes. They
enter a second host, which is swallowed by the final Vertebrate host in
which they become mature.
The Cestodes or Tape-worms have undergone more profound modifications both
in structure and in mode of development. They are all endoparasitic, and,
with one exception (_Archigetes_), attain maturity solely within the
alimentary canal of Vertebrates. In length they range from a few
millimetres to several metres, but this great size is attained from the
need for the rapid production and accumulation of enormous numbers of eggs.
The "head" or "scolex" is attached to the mucous membrane of the host by
suckers or hooks, but there is no mouth nor any certain trace of a
digestive tract at any stage of the life-history of Cestodes. For
nourishment they absorb, through the skin, the previously-digested food (of
the host) that bathes them. In a few Cestodes the body is simple and not
divided into "proglottides" or generative segments, but in most cases it is
jointed in such a way that the last segment is the oldest, and each
contains a set of reproductive organs. The life-histories of Cestodes are
most remarkable. The proglottides containing the eggs pass out of the final
host along with the faeces and enter the intermediate host with the food.
The larvae hatch, and boring their way into the blood-vessels, are carried
by the circulation to various internal organs. Here they usually become
"bladder-worms," and develop the "head" of the future sexual form. Then,
if, as is usually the case, the intermediate host is preyed upon by the
final host, the larval Cestodes enter the alimentary canal of the latter.
The head of the larva alone survives digestion, and from it the mature worm
is formed.
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