The Cambridge natural history, Vol. 04 (of 10)Warburton, Cecil
Science
The Cambridge natural history, Vol. 04 (of 10)
Warburton, Cecil
Animals
It has, therefore, been suggested by these authors that the
classification of the Malacostraca should be revised, and Calman (_loc.
cit._) has brought forward the following scheme:—
The division PERACARIDA, including the Eucopiidae, Lophogastridae, and
Mysidae (= Mysidacea), the Cumacea, Isopoda, and Amphipoda, is
characterised by the fact that when a carapace is present it leaves at
least four of the thoracic segments free and uncoalesced: by the
presence of a brood-pouch formed from the oostegites on the thoracic
limbs of the female: by the elongated heart: by the few and simple
hepatic caeca: by the filiform spermatozoa: and by the direct method of
development without a complicated larval metamorphosis. The biting face
of the mandible has a movable joint, the “lacinia mobilis.”[89]
The division EUCARIDA, on the other hand, including the Euphausiidae and
the Decapoda, shows the converse of these characters. The carapace
coalesces with all the thoracic segments, there is never a brood-pouch
formed from oostegites, the hepatic caeca are much ramified, the heart
is short, the spermatozoa are spherical with radiating pseudopodia, the
development is indirect with a complicated metamorphosis, and the
mandible is without a lacinia mobilis.
Corresponding divisions are made by Calman to receive the other
Malacostraca, namely, the PHYLLOCARIDA for _Nebalia_, the SYNCARIDA for
_Anaspides_, and the HOPLOCARIDA for the Stomatopoda or Squillidae.
The important array of characters which separates the Euphausiidae from
the other Schizopods and unites them with the Decapoda can no longer be
neglected, and the consideration of _Anaspides_ and its allies will
further emphasise the extreme difficulty of retaining the Schizopoda as
a natural group. In the sequel Calman’s proposed scheme will be adopted.
DIVISION 1. SYNCARIDA.
There is no carapace, and all the eight thoracic segments may be free
and distinct. Eyes may be pedunculate or sessile. The mandible is
without a lacinia mobilis. There is no brood-pouch, the eggs being
deposited and hidden after fertilisation. The spermatozoa are filiform,
the hepatic caeca very numerous, and the heart tubular and elongated,
with ostia only in one place in the anterior thoracic region. The
auditory organ is at the base of the first antennae.
=Order. Anaspidacea.=
=Fam. 1. Anaspididae.=—The mountain-shrimp of Tasmania, _Anaspides
tasmaniae_, was first described by Thomson[90] in 1893 from specimens
taken in a little pool near the summit of Mount Wellington; it was
redescribed by Calman,[91] who drew attention to its remarkable
resemblance to certain Carboniferous fossils of Europe and N. America
(_Gampsonyx_, _Palaeocaris_, etc.).
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